presented
    in table
    17,2. l4ean
    movemerrt rJistances
    r!irJ
    nol
    rtif
    fcr
    between
    arou0s
    .J -
    _r
    _
    of
    DI"
    U9l.vl999Jlrg.le-0ssd-J!.*[e-l!ed
    i
    nd i
    v i dua'l_s_
    f-Q.Ie_0Sed_.1*n
    .tf.qa!._-9..0-lnpql_e,-q.
    !o
    !ran5
    !t !qna
    l
    areas
    (t
    ..ri.?5,
    P>0.50)'
    but
    a
    signiftcantly
    greater
    averaqe
    rlispracemerrL
    from
    heit.rr.-r r{-)lc,l3e
    sites
    was
    _JLg!.e!.tgd
    detecte
    in
    .c.qrypirlson
    wit.h
    anbient.
    1bfent
    rel€i,.,c
    rele,
    sites
    (t=3.40,
    p.:0.01
    i,
    The
    f
    luctuating
    and
    occasional
    ly
    extreme
    temp_erature
    cycle
    which
    is
    characteristic
    of the
    heated
    location
    may
    in
    part
    contribute to a
    greater
    between-season
    displacement
    annng bass
    since
    power
    plant
    operation
    practices
    could
    contribute
    to
    considerable
    between-season
    variabi l.ity
    in
    water
    temperaturesr
    a factor
    which
    would
    rjemand
    relocation
    of fishes in
    extrerne
    cases.
    The temperature
    cycle
    of
    the ambient
    area
    <lf
    Coffeen
    Lake woulrl be
    expected
    to
    be
    more
    stable
    anrt
    preclictable
    irt
    the
    sense
    that
    only atnospheric
    events
    contribute
    to the
    ternperature
    variability;
    that
    area
    thus
    r.epresents
    rhe
    type
    of
    therrnal
    regime
    that
    northern
    largernouilr
    bass
    have adapted
    to.
    Distances
    traveled
    by
    markeC fishes
    were
    not related
    to
    either
    size
    at release
    or to
    number
    of days
    at iiberty
    as
    judqerl
    by
    a lack
    of
    correlation
    bclween
    those
    variables
    and respective
    displacement
    distances
    (Tafrle
    l/.3).
    Anong
    those
    f ishes recaptured
    dttrinq
    tlre f
    al
    I
    or sprirrg
    seasons
    followinq
    tlreir
    relcase
    (
    long-tenn
    movernenL
    s
    )
    ,
    _a
    !end9n.9y.Jqjeqrd-n._.r1il-h
    j-n,_q.
    ]:CS_t1_-cled_.dr.eas
    .was.
    -
    s'.19_e!!,94
    bV
    ![g:bt.gl"
    l-tg_g-ugtgy_.gf__f-qc--Ap-!ures
    rrlig.h
    occur.r-ed
    at or
    ne(rr
    origirral
    j*a!-e--iiJes.
    Approximately
    60%
    of long-term
    recaptures
    u,e,-e
    founO
    withi;-*
    I,000
    meters
    of
    their
    orjginal
    release
    sit.es
    anrl
    over
    half
    of
    that
    nurnb(:)r
    were
    found
    within
    200 meters
    of
    that
    location
    (Fig.
    I i.l
    ).
    DisplacemcrrL
    distances
    progressively
    decreased
    jn
    frequency
    up
    to.r
    maxjtnurn
    of
    6,246 rneters
    ivhich
    was
    the
    greatest
    displacement
    observed
    hy
    an individual
    aiter
    a long-terrn
    I)eriod
    cf
    I iberation.
    Crossing
    of
    the
    main
    lake
    channel
    by'largemouth
    bass
    was
    cornmonly
    observerl
    among both
    short-term
    and long-term groups;
    those
    crossings
    were
    det.ected
    in
    29i-
    afld 36%,' respectively'
    of the
    tota'l
    number
    of observations.
    Passages
    grrough
    the
    railroad causeway
    culverts
    were
    5% and
    93
    for
    the
    short-terrn
    and long-terrn
    groups,
    respectively,
    with north to
    south
    and'south
    to
    north
    movernents
    about
    evenly-,'djstributed.
    The najority
    of
    those
    movemerrts
    were
    observed
    arnonq
    individuals
    original ly
    captured
    and
    released
    near
    the
    structure.
    I
    T
    t
    I
    t
    I
    t
    I
    I
    T
    I
    t
    I
    t
    t
    t
    :1
    +
    '
    I
    i...,
    '
    ,,,,1:
    .'t
    I
    17.6

    .--l-'i
    -a
    ",a;t.:'
    a
    a::.::.
    !1!.ia:i
    =
    Table
    I
    7..:2, flegs
    anrl ranges
    of
    dlstances
    travele d
    by
    taggerl
    l6rgcmout!r
    bass
    Coffeen
    Lake.
    Rele.ue
    sit*s,
    sarnple
    size
    (ilj;
    dnd days
    elapsed:
    between
    release
    and
    recapture
    lmeins
    and
    ranrjes)
    are
    given.'
    Al'l
    observations
    represent
    long-term movements
    (iee
    text).
    ln
    Re
    lease
    Site
    mean
    'Says
    el,apsed
    range
    ,
    0is!gnce
    trdVeled
    (meters)
    n]e
    dn
    range
    He ated
    .,
    53
    Transitiona'l
    ?f,
    Anbient
    '
    46
    Lake-wi rle
    125
    310
    319
    297
    307
    I
    574
    147
    4
    671
    1221
    l4l-703
    I
    53-705
    I 48-690
    14t-705
    0-6246
    0-
    51 /9
    a-327
    5
    0-6246
    Tabl
    e
    I 7.3.
    Correl
    ations between
    distanced
    traveied
    (Log
    [meters+l
    ])
    by
    tagged
    largemoutlr bass
    and
    selected
    independerrt
    variables.
    Separate
    correlations
    for short-term
    and
    long-term
    movements
    were
    rnade.
    Sample
    size
    (N),
    correlatlon
    coeificierrt
    (r),.
    Students
    t-value,
    an'd, significance
    probabi
    I
    jty
    (p)
    arri
    qiuen.
    Independent
    vari able
    Short
    term
    ,
    -AJ_D'avs.
    e I
    apsed
    B) Days,
    el apsed
    Lonq
    term
    -AFTTI
    release
    B) TL at
    release
    .l54
    12s
    'l
    s4
    125
    -0.r7
    -0.01
    -0.00
    0.03
    0.01
    >0.
    50
    0.07 >0.50
    0.01'
    >0.50
    0.32
    >0.50
    .::..:.:.
    f:...:.
    -:-
    I::.,:
    -:':.:
    ''':
    ':
    '
    '
    ,:.--
    .1';t,",
    ii
    !
    :.n
    17
    .7

    i:..,:-,.'
    It
    ):'.'.
    l'
    l,
    I
    I
    l'
    :
    t
    I
    t
    T
    I'
    l'
    I
    l'
    I
    l'
    lt'
    FggsgF
    EFsgss
    'rltaa
    oo
    6O
    crO
    a-
    o.
    o
    GI
    %
    Distance traveled
    (
    m')
    from'release
    site
    meters
    )
    from
    rel
    ease
    s
    i
    Les
    in
    Coffeen
    Lake,
    Al
    I
    taggert
    recaptured
    during
    r.he
    foilowing
    Fig.'.17,1.
    Frg,l:,g1.{
    of displacement
    (ranges
    in
    exhibited
    by
    tagged
    largemouth"bass,
    fish
    were
    released
    in
    the
    spring
    and
    fa
    I
    I
    or
    spr
    i
    ng
    seasons
    .
    l,i
    I,,
    N=125
    1.7.8

    ;',
    :,',,:::
    ,
    :,;
    j..:..
    -a:.,::.
    :
    :;:.,,4_-:'
    :.
    :::.:i
    ;;,,-t:,;t
    ;":',,,.,,,:'.
    ,l:,,,
    .1,t.''.
    t',
    :I.::
    '-
    --.
    .
    .
    :.
    :
    ,.,,
    .,;-'
    ,;,.
    ;'--;
    ..
    :I
    ::
    i..::
    t;l
    ,:
    I.::
    'I
    .:'
    -.
    I
    :,:.
    I
    I
    I:
    .:
    :I
    I
    I
    'Knowledge of the
    life history
    of
    larqcrnouth
    bassris
    exLensive
    as
    eviderrceC by
    a
    .]
    :
    ',"large
    body
    of I
    lterature'r(Robbins
    antJ
    l'lacCrirnncn
    1974,
    St.rorld
    and Clrrt-rel'
    .I975,
    '
    :...:
    '
    rnovernent
    Heidinger
    patterns
    1976)
    but,
    of
    tltis,
    until
    spec
    recently,
    jes.
    Coutant
    I
    itile wa::knswn:cf
    (1975)
    rnairlLaine('
    Lhe
    dail.y
    Lhat wate!'
    o.r
    :cdsilnal
    .
    '::
    temperature
    was
    the
    most important
    f,tctor
    governing
    bass
    Inove,nents,
    dr'rit
    ,
    in
    'ndture,
    individuals
    adjust
    their
    positicn
    cieily
    and
    se,tsr)nal ly in an ef
    fort
    Lo
    locate preferrert
    temperatures.
    Beyonct
    thaL
    basic
    behaviorol response,
    t,{o
    .', general
    i
    zed
    patterns
    of movement
    e;nerge
    f rorn
    the
    I
    i
    teratrrre. Sorne
    invest
    igal'or-s
    'have
    noted
    that
    bass
    movement.s
    were
    concentrated
    within
    a
    well-<Jefined ared,,
    an
    ,,:observation
    which
    led to
    the
    concept
    of
    "home
    ranqe"
    {l-ewis
    and Fl
    ickinger'
    1967,
    t'Jarden
    and
    Lorio
    1975).
    Ihe
    location
    of such
    centers
    of act
    ivity
    wer'e bel
    ievr:d
    to
    change
    seasonal
    ly. Fetterol
    f
    (
    1952)
    offered
    tlre
    viewpoint
    that di
    fferinq
    ,,'.
    innate
    characteristics
    among
    in<iivicJual
    bass
    may
    troduce
    an extensive
    rnollility
    in
    some
    dnd a
    more
    sedentary
    nature in
    others.
    Distances tr.avrlled
    b.v
    bass
    in
    -
    Coffeen
    Lake
    (ranging
    up
    to 7 ki lorneters)
    were
    rrot
    exceptional
    cornparerJ Lo
    oftret
    studies reported
    in the
    literature
    since
    the
    rnagnitude
    of
    qlovcrnenL
    exlribiter!
    by
    this
    species
    appears
    to be
    lirnited
    only by Lhe
    physical
    lirnits
    of
    its
    environment
    (Heidinger
    1976,
    Tranqui
    I I i
    el;
    al.
    1979b,
    and
    oLhers). ExLensivri
    Oisptacement
    between
    seasons
    appat'ent1y
    was the
    exception ratlrcr
    Llran
    the
    rulcl'
    in
    Coffeerr
    Lake since
    ntost
    recapturerJ
    indivirJuals
    werc
    fourrd
    within Lhe
    srrrne
    gelteral
    area during
    spring
    and
    fall seasons.
    H0wever,
    the frrlrlrrerlcy
    anrl
    magnitttde of
    tnovernet'lts
    which
    occurrecl
    drrring
    the
    interin
    periorl
    cor.rld not be
    determined
    with certainty,
    and the broa<i
    range
    of dis;,1acenenl.
    rlisLancr:s
    I
    ..
    '
    observed
    after
    oniy a short
    period
    of liberty
    suqgests
    that Lhose
    r!istances
    may
    have been
    extensive.
    Coo'ling lakes
    provide
    a rather
    uniqtre envjrorrrnent for
    evaluat,ing
    fish rnovc,nenls
    since widely,livergent ternperatures
    gerterally yrersist.
    In unheaLerl
    lakes,
    thermoselective
    move/nerlts
    are
    usrral ly
    1lrni tetl Lo
    vert
    ical
    arljusLrnerits,
    whereas
    cooling lakes offer
    a
    rnode,
    of
    thermoselective
    rnovernenL
    jn
    the horizontal
    direction
    as tvel l. In Lake
    Sangchris, Tranquilli
    eL
    al
    (1979a,
    1979b)
    f
    r-rLrnrl
    that
    averaqe
    L+--"_
    daily
    movetnen!s a$ gbso-l-ute rlistances
    !1av.aled
    by largernouilr
    Ilass
    ,werq-lilqilicantly
    greater
    amonq incliv i<Juals
    orillirlal
    ly capturecl
    anrl
    releaserl
    jrr
    ,------*--:-
    -:__-._..-..-*
    .
    .: ..
    :.
    .
    :..
    '
    -
    llreated
    areas,
    a
    finding
    which was supporterl
    by tlre results
    of
    ttris
    sturly. The
    authors
    predicted
    a seasona'l
    cycle of
    attraction
    and
    avoidance
    at Lher,nally
    extreme
    locations, whereas
    l'ishes
    inhabit,inq
    therma)
    LransiLion
    areas
    wcre
    I
    I
    .:
    I
    :
    ,I
    I
    I
    I
    I
    I
    !
    I
    'l
    l"
    17.9

    ::...j
    r.:ta
    =l:..:.i:
    't':',.::1
    .
    ,
    :.n.:,: .
    i;i:'.
    ::.'i;i.',
    :.::::
    E,''
    ..:;,1;"
    t;:
    .,:t,
    -i:,4::
    ;
    .
    ;,:..
    ,
    j:";
    :
    ji:.:.-
    :
    i:l:.
    .
    '
    .:
    ::.:
    .
    ;'::':
    ,
    :':i,:
    j
    i;:,..
    ?::t,,.
    ':::
    :-
    !:..'
    .',..,1-
    '
    li
    :,::
    ,;a'.
    l''
    I','
    believed to
    be
    most'sedentary.
    ln contrnst,
    Qrrinn
    et al.
    {l9iB}:conclurjed
    fro,n
    their
    study, and
    earl
    ier
    studies
    of
    bass
    movernenLs
    itr
    ,l
    s0{rLitgasLrlrrr co0I
    in(l
    reservoir,
    that
    f
    ishes in
    heaterJ
    areas
    r{ere general
    ly
    most sedcntary arrci
    exchante
    of
    individuals between
    heatert
    dnd anbieni,rreas
    vyds
    {rnc'drrmr)n.
    The
    fishes
    observed rrere
    found
    to trovel
    extensively rlr,ithin the lahe
    even
    thouqh
    their
    movements
    were dpparently
    unreldted to lhe therrnal
    graiJient.
    Population
    Estimates
    A total
    ol'
    I
    ,1
    24r
    l argernouth bass wrtre
    captured by
    el ectroshocking
    rlrtr ing
    the
    1979
    populati0n
    census.
    Inclurled *ere
    271
    indiviriuals
    (2lSO
    nrm
    TL) that
    comprised the recapttJre
    sdmple required
    for
    population
    estimation.
    0f that
    number,
    54
    had
    been
    previously
    nrarked.
    Therefore, the
    estimatcd
    popu'lation
    size
    as of May
    1979
    was
    4,958
    jndividuals
    for the
    size
    group
    considered
    (Tahle
    l/.4).
    The
    951
    confidence
    interval
    for that
    estimate
    ranged frorn'3,734:to
    6,132
    incJividuals.
    The l9B0
    census
    produced
    1,124
    inCividuals
    includ'ing
    340
    (>360
    mrn
    Tl-)
    which were selected
    for inclusion
    irr the
    recapture
    sarnple. 0f that
    group,
    65 had heen
    previously
    marked
    yielding
    an estirnate
    of 5,466
    inrlividuals
    and
    a
    95X confidence
    interval
    frorn
    4,285
    Lo
    6,647
    (Table
    l/.4).
    Ilre
    two
    estirnates
    were
    in
    close agreement over
    the two
    years.
    Co11elp.ol{US,__s!1.11l1rLg_gfg1)
    estimates, 9.3
    and 10,q,_E/ha
    (Table
    17.4)
    were
    similar to the
    l0
    kg/ha figure
    'eTUEa-WAlenkins
    (.|975)
    as an
    average for
    rnid-southerrl
    reservoirs,
    arul
    to
    Llre
    Lake
    san
    gchr
    it!
    _q4+sI€j[*UJ]:
    estirnates of
    B,B
    kg/ha
    and,l2.6
    kg/ha,
    -jg:qg$_Uety_
    (Tranquil
    l'i et al. 1979b). Estimation of the biomass
    of
    largernoutlr
    bass
    by this metlrod
    yieltled
    slightly higher values than the
    7.7 kg/ha
    estirnate
    rterived
    by
    cove
    rotenone sampling
    in
    Coffeen Lake
    (section
    14,
    frerein).
    The
    latter
    techrrique
    generally
    yielded
    few
    individuals
    of
    largi
    size cornpared
    to
    catches
    obtained
    by electrofishing. The mark anrl
    recapt'ure
    estimates excluded
    individuals
    (200:mm
    total
    length and
    thus
    vJould have been
    sonreirhat:higlrer had
    they been included.
    :
    l
    Fi
    sherman
    explgr!.!.at
    jgn,
    ,
    ,
    From the spring
    of
    1979
    to the
    spring of lg8l sport
    fishernen'reportetl
    B0
    catches
    of
    tagged
    bass and
    84
    catches of untagged fish.
    0f
    the total,
    90 fish
    weq"e.
    kep!,:,28-'t{e,r:e,,released,
    and
    the
    f
    ate of 46 was not reported.
    The
    shortest
    I
    I
    T
    l,
    .
    l'
    t',
    t
    T.
    I
    I
    I
    .
    l'
    '
    I
    I
    ',,.
    '
    ,
    ,.
    '
    '
    '
    :a,t-,:
    .'-.',:a::. a.
    ::::1:
    I
    7.10

    Table
    :
    '
    17.4,'.
    '
    ''
    of
    Hark
    .rarseryourh:
    and
    re99o!ure
    bass
    22'00
    populatlon
    *
    iotir
    cstirriate
    i;;;i;
    s
    l;
    (r
    ir'r'1;y'-ibll-u,'a
    ?
    st:anrlard':errors)
    'ruy
    average
    we
    i
    ght
    ,
    and
    s tanrl i rrg
    c
    iop-
    u.e
    -g
    i
    ven'.
    Parameter
    Ita
    I
    ue
    Estimate
    l
    i{umtrer/h
    a,
    Av
    e
    r'age
    we i
    gh
    t
    Standing
    crop
    Est imate
    l{umber
    /ha
    Average
    weight
    Standi
    ng
    crop
    I l.l
    0.840
    k9
    .
    ,,
    g.,kq/ha
    l980lopulat
    ion
    Est
    imate
    546rr
    t
    I lBl
    ,
    l?.3
    ,0.
    BB2
    k0
    l0.B
    kg/lra
    | 7'.1
    |

    '.::
    aa;'
    '.
    i
    :.
    :'
    .
    t
    a
    ",
    t ime
    i
    rltervcrl
    betreen
    tagging
    and rep0rted
    capt_ure
    by a
    f
    i
    sherrnart w,ts
    J
    ,j,lys,
    the
    longest
    6S9
    rlays.
    Sixty-two
    successful
    fishing
    rlays
    werc
    rep0rtcd,
    I
    in
    February,
    5 in
    l'larclr,
    6
    in
    Aprl l,
    zB in
    l4ay,
    l5
    in .lr.rnr',
    4 irr
    Jrrly, z
    in
    llovernber,
    alt,C
    I
    in
    Oecember.
    Ihirty-nine
    partics.r;iire
    reporLutlly
    fishi,rg
    th;/
    'south
    part
    of the
    lake
    (soutlr
    of railroad)
    and
    seveft rlrer*l
    f
    ishing the
    nrirLh
    f>
    \
    part.
    ^-/
    Al
    I
    of our
    catch
    records
    were
    obtained ilrrough
    volunt-ary
    reports,
    usudl
    ly
    by
    telephone.
    ,
    Since reports
    were
    only received
    from
    fishermen
    who were successful
    ,in
    catching
    a tagged
    f
    ish,
    we cannot
    quantify
    the rate
    or
    impact
    of f
    isherrnar't
    exploitation
    on the
    bass
    population
    other
    than
    that
    which
    is
    inrplied
    by
    thr,.
    above
    rtata.
    At most,
    exploitation
    of this
    species is
    rnoderaLe
    cornpared
    to
    public
    fishinq
    lakes
    (Coffeen
    Lake
    is
    closed to the
    public)
    and the
    reader
    is
    rerninded
    that
    population
    dynarnics
    of
    the
    bass
    population,
    as
    re')orLecl
    herein
    and
    in
    other Sections
    of
    this report,
    Inay
    in
    part
    reflect
    a
    cornparatively low
    rate
    of
    fishing rnortality
    relative
    to
    oLherr
    Inore
    heavily
    fisherl
    la<cs,
    f'
    ''.
    -l
    t,
    T
    T
    !
    T
    I
    t
    I
    I
    I
    I
    I
    l'
    I
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    l.'
    t::
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    ':.
    :: :
    l7
    .12

    :..:
    ::''
    i:-
    ,,=::
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    '
    '
    l
    l'-
    . !
    r-RA:i.,iti
    .
    l
    :.
    .I:
    -.
    I
    Coutant,
    C,
    C.
    lgTS,
    Responses
    ,)f
    ir.rs5 !rr
    nr:,;'::
    ,lr,,i:
    rrt:!':t::1
    :r.,:'"..
    r.'.
    rr+-
    '
    ,.
    regimeS,
    Page
    S
    27?-ZA5
    f
    n
    A.
    l{.
    -!trsurj
    rnrj
    i.i
    .
    :.;q;;grr:r-.
    ,.,i.
    .,
    ",
    ,
    ..
    ..!.
    ,
    , ,biology
    and
    rnanagenent.
    Jport
    f
    i;hirii'lnst.if:;i,-', i1'ri1
    i..;1
    :'r
    [Jell
    ,
    M,
    g.
    1958. A new
    fish
    trg
    arrrl
    rapirJ
    c,trl,"irj1,.-i,',i
    !:.,ir,.i..]',
    .'.
    .i
    :'-=r,'t.
    Am. Fish,
    5oc.
    9/:5?-59
    Fetterol
    f
    ,
    C'. dr: la
    llesa,
    Jr.
    1952.
    r1
    pr:pulitirlr
    l,lintergreen
    Lake,
    l'.,tlarn,tzoo
    Couni;rr
    :4i{h1rinn.
    effect
    of netlinq
    on
    coildition.
    :,1.5.
    Ihe5iE,
    127
    prt,
    \r
    rj:i
    j
    r:t
    !-
    it:
    :
    f
    i.r!lr":.
    :-.!
    f
    iit:n
    n'.'i;',
    .i!'',ir-i',t,r.
    t',1
    '.'1
    'i
    i
    q
    i
    l
    grr
    :.i
    4,
    ,-
    ii;
    1
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    ;';,r
    .
    ,l'
    ' ,.t
    .'1
    ,
    l
    I
    ,],,
    .
    :!::
    ,I
    I
    I
    Heidinger,
    R,
    C.
    l9/5.
    Life
    hi.itcry
    dnrl
    l:i17
    li;1;
    ,;f
    i.nq
    )r'-.r'],',,,i
    !.,r
    :,:.',
    ll -?0
    ln
    R.
    H.
    Stroud o;td
    l{.
    Clepper,
    e{|s..
    1l*ck
    br';r
    rrjt',ii.:.1;r
    ,r;,!
    manageiiEnt.
    Sport Fishinl
    Ingtit,lte,
    ';{,!stii,r'1t
    lr, f
    .i..
    i._t:jri;
    ;T
    .:a
    rl
    Heidinger,
    R,
    C,
    '1976,
    Synopsis
    of brri!oqii:a]
    ;!
    rr-,r
    ric
    1i!i:
    :
    i.ii')r'::i
    i'it
    l-'1,i,
    ljlqropterus:g]gtdE
    (t-acepecie),
    1,ji)?.
    i,1{i
    irsiri:r-ii:r::r:j:i:.!,}..
    !i-i.
    iir,
    6f
    pp----
    -
    Jenkins,
    R.
    !4.
    1975,
    Bl
    ack batt
    crops
    in(t sili:cir-i!'.!\ir,1,..!!
    !
    r.,r
    !ri
    r,,,i,.;'"'r-:r..
    ,
    Pages
    ll4-l24
    In
    ll
    .
    H.
    lrtro*rl'an'J
    l.l
    . [ie;lper,
    r.d,,..
    iil
    i-'..
    lir.,'-
    yit,1=;tt-a.r
    ,iz;.2
    .
    tnanagefient.
    Sprx"t
    Fishinq
    Jnslitute,
    *'dshirlltrirr,
    :i.il .
    l
    Lewis,
    l'l
    .
    i,l
    .,
    Jr. and S.
    Flickirrger.
    i96/.
    ii'arD .rlrr:i!'.r',t1,t,,
    t
    tt
    rf:r,
    large'routh
    bass
    (t'licropter:f
    ::]p_:4e.f].
    icr'i4t1v
    i,1.,,ii;!:1,:rj-l,i;J.
    ,.
    Quinn'T.,G.t.,.Esch,T.C.Hagen,anr1').H.sitlhonl,|91v,i.tj|t-j'-ji!|=:il.
    i,ItroV€r0rlflt
    and homin!) by I argemouLh
    bas<
    tl,ticroplerus
    1,
    llsv;irdr111
    :rr
    i
    ::therinal1yalteredreservoir.Copeia{r)5{"*.-Trr:
    Rot1ers,R.A:.l9B0.FYl9iB.:Iliinoissp'r|'fi.;hery\'ir'/!j''!p.l'.1ii.:*:'-ii'
    Report
    No.
    50, Illinois
    Dept.
    of
    Ccnser'ytl
    irir),
    0ivi,;ior
    ,.ri
    l-i',ire''r':,
    ,
    Springfield.
    Rohbins,
    l.|.
    lJ.
    and
    ll
    . R. l,lacCrimrnort. 1g74.
    tlrt-'
    bla;kl;.tt,.
    ,;{
    ,'r,':r1.,r
    r{
    I
    ,i,r'l
    ,
    ,
    overseas. Siomanagentent and Research
    EnterprisitE,
    llrrt-rrlr'r
    irtn4*i.
    !!6
    pF
    r
    Stroud, R. H. anrt H.
    Clepper,
    eds,
    1975,
    Bl,ick
    basirri(il()'ly
    ,1.,1
    ,1i-.t;,,,!,rt'.
    Sport
    Fishing
    InstjLutc,.'Iashirrgton,
    D.C,
    53/r
    pp.
    Trartquill
    i,
    J.
    A., D.
    W. Duf
    ford, and R,
    l.l.
    Lariroore.
    197'1.t.
    lit
    iv:',,rr',1,i,:
    .
    preferred
    terrperature
    of
    radiotaqged
    larrlernorrt,h
    llas.; in
    l.ik;'',.1r1'llliri'.;,.lnri
    Lake Sholhyville. In Evaluation'of a
    cooling
    l,rke {r.;nery,
    v,rl,
    :i!,
    Il!.
    :'
    Nat. Hist,
    Survey'fln-al report to
    Electric
    Power
    Rrlr,ear(h
    !nqt
    rl-rlte, i'dit
    Al to, CA.

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    ;r.tri
    t
    ti
    r.
    jf'rf
    1
    l!
    1,)rr
    iiui. r,:i.il
    ?
    rr;.
    r;;-.1:,,.
    |
    .
    t
    .r.,
    ,r.'1,
    .::,
    ,.,.,
    .t,.:
    i.,r,
    otf
    rl;rt
    {itr
    :;i
    /glr!,;r.;.-r11
    trJi;iii
    .;i
    lrlr-,
    irilr.,r
    |
    1\r}
    r.t.
    ,-
    ;:
    ).,, I
    ,i
    I
    .,,
    I
    i:.:,,
    ftil4ti,inSrtip
    nr'?dfr,rt h,r!r
    i::!!rj!'.i,;.]
    !.tiJ
    :ri!,,,ri.
    t.
    1
    i.1t.,..!
    )i
    r,r,,
    .1:.-.
    -.rf
    t*f r,:fii+l
    +f ?fitf
    ilf
    ;ijrif{+e
    =i
    t-+1.!l:
    1q:..:;1is,
    i;;,t1
    .:rt,.
    '
    rt
    ,:,
    tt,:t.,
    .;
    5iJtri(ielrt!.
    iitlllt:
    tS.]i
    +,ri it
    rl!'<-'1,-=1,0
    irr
    E)-!i/:it,'..,....
    l:
    i.irt,:i
    .
    ii,
    ,r
    t*b,/l,.lt13{i
    r-*JlrJ*
    ,lt'
    i:,ll},
    ,ti!.';r
    ?*
    3t:1,;,:,t:.,,
    ::i
    !'1..,.,sj.;,q:r
    i i.j,-ir..1;:,,,i::.
    ti'J.1
    if
    ilarre+
    prtb,thiiity
    r3f
    p..-{}.i}f.
    i,i:'::,r
    !ti'
    r,i:a
    rilr.:
    j
    r
    -.'rf i:,
    r:.
    ,-,.
    iIaii:*l4..}},ahi3htorfij]t!1i:1r.ri5.1l1;;l3fiii..j:1Jir€!i]i}rl.]i:i'.ll:,l-t,.
    c,ilrrif
    ie
    trtt.
    Ihur,
    int*n:,rty rlrf
    r),tr,ltir,it
    ititt.,.,..ir
    r.1,,,:,
    i,l
    ir
    j.j.,rJr!i,,
    -,,.;t
    L{tnLt
    ilrufe
    to a rltot
    i'te{y
    }a"
    i,url1
    {*.j!1it
    l.;
    iqri ::;
    lil,;
    [,.5,
    :r,1r
    +ril
    r 3,tr?
    ,
    Lhg
    *bserved
    variatian
    irt
    q,OndiLiOrr
    rar
    rl(.i.r-r:rri?i..;
    f
    .rr
    !,.;
    !ilr:'i..:,_i'ri1=1::r!i
    Iraf
    ds
    i te
    dens
    i
    ty.
    :.
    .1:."
    I
    .'j
    :
    :!
    ;l
    j:-:
    ;1!-t:'
    il
    Results
    of other
    ll"udies
    Suqg+gL
    lh+t l.ar:
    lrr!eri:,
    i:..,1
    ;i!
    l.
    .:,!r1
    l:,,!;1n
    iilt.,,:.
    1.11
    r.,,:r.
    oh$Crv€,t
    in
    Cof
    f
    ccn
    Lak4
    l)lxi-'gilit
    ,,ray
    4rjrrr,r.
    ll-*r;t,,
    t-
    i
    r,1t.ir.
    ,i,:a
    .p.,rir,.t;
    ,.r?,-.
    gnlthernarl
    (l96tt)
    tesled
    thlr:,t
    lflv{.tli
    af
    rof
    ,icl,1l
    r;i;1.,
    lrr,!
    f
    ,r,llr;l
    !
    ri4r,
    7rt
    I
    .,.jr..i.j.:
    total
    ddosity
    of
    10i.3
    trct,!cdrcdf
    i,t
    pr-rr
    risrr
    iJ,).i
    r11
    ?.fi..
    liv,:r.
    j
    r!:,!
    ,,,:r
    i,i4.,,-ri
    Jrowth
    of
    Infecterl
    hlueqills
    Cornparcr!
    L,.r
    .i
    ,:r)rrlr..r'i
    .;r,i,1gp,
    i:.,,:
    !.r,,,..,
    1,r.,
    il
    j.;i.
    I
    (.112.5
    io
    liver tissuei)
    r'lnrt
    546.g
    i?d|J.1
    in
    l:v,.r.,i
    ii,rr..rrit,..,
    ,,i:1rr;!
    r,,r
    riiy
    f
    edu(',ed
    {rogtlr
    of
    :
    lllfectf;rJ
    vl:rsul
    cr4tr.1!
    rJrirtrlr,).
    ii
    :_
    r,r:r1h
    t'rr:
    !,_,.;,:I
    ;
    rt
    t4.3

    --!..:,,.,..a.
    :.:t:
    .
    . ,:.''
    -:'::
    ,:
    ai..
    ::l
    ''a::::,'
    ..::
    :;
    ,:.
    i:.
    :
    rt:i:,.
    a:...'
    ..
    '::.
    l
    .-,
    '
    ::::a.,.
    '
    ,'
    ,.,
    :::,:'
    ..
    ::::
    l
    :;:.1
    '
    -
    ::::::
    .a:
    :
    i,:'
    :'i
    tafrle
    18.,1,
    llFlia
    tiir:l;t..
    +{
    1'. E.:-{rlr.:rr"
    t.F\":,
    {.,,:iti,i
    .',i:
    i
    iir!
    'z
    '.}l ,.':tli{:,':at
    ,i,lt.:
    *1u*€il
    ig.-'
    :i-
    11?.= lr;"..:!
    i,;4t
    ",.,5:i:5
    1,,.
    ',
    t
    1;'
    :,'i,,
    r
    t',;t''.
    it-"lrle-:ilt
    L-tl!:,:i:,
    4
    i}
    ":!grrtf
    il.1.::,*
    ;ir'::i::llr!::!
    t
    iir:
    1r-i.
    j',"
    {hq
    t:-trlg,t
    i:.i*Air'i
    ++l
    *;ii
    i;ir!'f
    ;ir'::.;:r:
    ,,:!
    ?
    t'
    :tt:'..('tr.:*:
    I
    ,i
    i!
    t.
    t
    i-ocat
    iorr
    _il.g:1:
    ji-qru:*g!."
    a,1i?1
    i'
    I
    :"+:l
    =:1*
    --'-"**:-.-:*'
    "1"-
    -
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    ar!
    r
    4'|iJtl
    t-
    l'
    t'
    l{e6t.1/t
    ;''nh
    i
    r:xt
    1t\1
    .7
    7!r\.i.1
    1
    q;+
    l /
    rj't
    i!'l
    4*
    :-
    .lr
    l.fhlrl
    18.? Girrei
    tl
    ifift
    y;lt"6e{,111
    rjir!!!i+r.
    .,
    *{ delf
    cCn Lakrt
    b
    1ari,;i
    !
    il
    ,
    ,ff
    the
    host-
    fi:h.
    '>-1ap!1
    qi
    von,
    r-il
    fi.
    +inirrit::i
    I.Y.;l;
    is;
    l!;i,r1' :.!5\'J!:5
    {
    ,r:i)fa'J{ff''Jiir,tll,l)
    !il.i i :i li
    1.1't,ll,'i
    iifr-,
    .|'il,
    r;
    ,-11'fi?'l
    4!.
    i'rrl
    r.
    it'l
    ttt'.ii"i?
    ii:;rri
    l'!{..inii..?
    pr'+ir'l1r!
    liiy f
    i;
    ,rr'tr
    I'
    I
    I
    I
    IrtrJepf
    nde
    ql
    l.
    .rarinillc
    Iot"al l{-'ngt,t
    9:i
    :'r i0
    l.i.:]I
    T.
    I
    I
    I.
    I,.
    l:
    Table
    13.1.
    Irart
    i,rl
    eorrelatiort
    clt':f
    f ic
    jiinL
    i!'/iiJ't;i
    t)':!'a{rlr'i,
    $tl:nlii;r
    +i
    P. rninilnrt'n cysIs
    in liv*r's
    of
    (,offrletrl
    i-.tki'lrlrtr'1
    ills.trtr!
    FeliTiTiTeights
    of
    hosf
    f
    isltr-'9
    *her.:
    A'nrtrh4r'
    r].l.,l5it'ts,
    B'
    relative
    *el[ht,,rnrl
    C=r-cLrl
    lprrgth
    {ni:}d
    eonst,ant}.
    Slrpir.'
    cire
    (il),
    der;rae5
    cf f.'le(to::r
    {C.f.},
    +nrl si1;tif
    icartcr:
    prolldtli
    I i
    ty
    (
    ft)
    are
    9i
    vt'n
    '
    Var i
    atll c
    d.
    f
    .
    rAlr'c
    Relative
    weighf
    9ft
    95
    lB.4
    -Q.ll
    ''1.'15
    t;
    ::...
    a=
    ::,:.=
    ,",,tj
    t
    a:
    .l
    a-:
    '1,.
    a.;,,
    ,1..'
    :i:
    a:.
    -:
    ::'
    ..
    ..:
    -.-:::
    t:::
    a
    :::-]::
    :;.i:
    j.:
    -:.:'
    t.a,,.:
    :.:.i: -,
    ',,4-
    -.-:
    iaal-:
    =..:.,:
    .i-.,
    i,
    ,t.t:.
    :.
    ,
    ::=.'

    ;,:Ir-.
    a:'I.':.
    '.
    ,.4:
    :,.
    ira:: j:1:
    =lt
    '.','-.j.
    i]
    :
    a..: ..
    ,l
    :...
    I
    in{gstdt icrt
    in Lhit
    st-'t,1!
    n.irrj drr't
    'f
    ic
    il
    l,
    :r:.-1
    .;,
    r:
    !,
    !
    r,,,
    I
    ,;. ,y
    ,,.,i
    i'
    ,, :,,',.
    :
    UlU5l,'iinnat.jr8dR.,lt{}in;}-lrl|}il-l.!f,!!1,iit*..,1,...1.lt,.,'11,,t.,lj.r.
    5tUr|y.L+wisanJliii..i:.rn{l;l6.1i.;llsi:rvd,''|}.1t'1r.}i1rii.,.i'?],r|..t
    trlUegilli
    rdn{)inl
    riit
    r-n
    l{}Jrr
    pnr'
    t
    rsn,
    fr,ri
    t4r
    ;
    {
    :.,!'.7
    .r
    ,.f
    i,,.
    |
    |
    f
    .1
    ;r,;;
    densltieS
    4li
    cnnrjitiolr
    cf
    lne h0sl
    t i1fle1.
    i-r,;i-rar.i:ril-!j
    ii{
    ii.
    ,r,
    lrii=;,;;';
    infertaLions
    I.li lfill:'
    in
    hii,i';illr f
    r0r1
    L.r..i-,
    1,;r1.1,;i,rir,
    ,
    ,,,,.i,.;l"il,i,-,.
    r,,
    crro l
    in1
    l,rkt,
    hut
    0lrl.i
    56{,
    tnrl
    0f t i4trL,r
    if
    .rt'.rtv.
    ,:
    n
    *,}il
    ,i
    1,n.,,.ti1-,,r
    il.:, r
    centrdl
    Illil'roil
    f
    117ritl
    ql1tLrDl
    rr.lrarv0ir^
    (il;r;r1
    !,i
    .!
    r,!
    lr,trr1,1i'1
    1, irii,:l;.
    i.
    i,,
    also
    notcxorthy
    Lha!
    grlivth
    rf age
    !
    an'i
    r:lder
    lliue,!ill.r
    w+l
    eliir;r'r-.1piri in
    ..{i.i!
    Shelbyvi
    llr: tit,tn irt
    Lakc
    i,rrr.1Chris,
    whicrr
    h,.is
    I
    ,,i
    rril,t,l
    nl,r",lr
    ll
    prl),rl1l i;rr;
    si,nilar
    ttt
    Lhot
    ql
    Colfeerr
    Lake.
    fleaSons
    -flr.Lhe-.pfOliferat
    ion
    of
    yrl|i i.g
    !ivt,r-
    ir,ih.,
    rr
    C,-r{'t.Jr:.rr l.,tki,
    tr-r:
    iir.i
    ly
    COnjeCturdl.
    3r'eat
    lllue
    hcroni,
    rrhiCh Servr,
    f,S
    t.ilr'
    ;:irirrr:ip.rl
    r!,tf
    i11
    ii
    jTc
    itr.,q!
    1.
    in
    the
    lif
    e
    6yg.le
    of
    this
    species, wer',1
    {)ccli,i,lnally
    rbsrir"v,.trl
    irr
    I
    itt,)r.li
    .jr',r
    i.,
    of tlre lake, huL thtjy
    vJere
    rtci.
    coin.llori.
    Tlt{,
    stettply rlrpinrt
    .,hrrrr?1i4,..
    6iri6ir
    i
    ,
    chardct,eristic
    of Cof f
    rlun
    Lake,
    I
    irniterl
    v.t:;['Jl rr
    pl,111t
    r;rorrth,
    t)r'irrr:r
    iiy
    tgJotogglg1}
    r,pp,,
    f0
    .t
    ndr'row
    llanrl
    par;l
    lnlinq ttr,,3
    5i16y',ili41:. illrrr-'r1i!i,,
    t.ypic.tl
    ly
    cortgreq,tte(l in
    thosr:
    areJs thr0rtrllro,tl
    (ndsi
    )t L,rs
    ,;r:;1ir!.J
    (,:
    !'j,rn.
    Corn'non
    rJSagC
    0f
    liLLor.tl ,trrlas
    lly
    rlef iniLive
    lroslg
    {it*r'lrrii
    .lrrJ
    s,-,f
    {),)'1
    internerli,tte
    ltosts
    (f
    ishcs)
    probabll
    Derpr'tlatr:s
    Llrt' I
    if
    r;
    q'r.lr:
    lf
    P.
    :jlll,,:,I,:
    :",
    Cof
    fcen
    Lake
    sitlce
    large
    nurnbers
    of
    blueqi
    lls
    crrncenLritlerl
    irr
    ri)st:r
    ictr:iJ
    shoreline areas w0ulqJ
    prrrvide
    a
    readily-,accessible
    fi;ragf basc for: pisciv,lr'r.ru5
    birds such
    as ltcrorts.
    As inrJicated,tbove,
    int.crrsity
    rf
    irrferLa),ions
    ()b.,,rrveri
    i:r
    Cof
    feen
    Lakc are certdiniy
    extrerne
    erioutllr to
    ret,rrd
    ,J'^orLh ,*rf
    lrluerli
    lls
    arrrl
    ,r,r/
    have
    contributed Io Lhe re]ar.ively
    low
    conrlil.ion
    r-rf
    larqer inrJivirtrrals,
    llowever, isolation
    of
    one
    cause
    for
    llrc
    sLunteqJ,
    n,tLl,'e
    of Ltre
    pcrpiri
    rLirtrl
    164y
    '
    leacJ
    t0 an
    errot'leous concluiion.
    0blter
    rrossible
    conllr:ibulirrll
    f.lctors,,Jisc,rssc,J
    elsewhcre
    in this report
    (Sections
    7
    arrd'15),
    irrclurJe
    n
    relJtively lo,a
    derrsiLy,-
    of
    'ljenthic (foraqe)
    or.ganisos,
    lack
    of
    Euf
    f
    icieni
    prcrlaLion
    Lo
    prevenL
    '-
    "'
    overcrowding of
    bluegi
    iis, ancl-cotnpetition
    f
    rorn other
    srrnf
    ish
    specir.ls
    rr'Li
    lizingi
    the''s
    aine,.f or
    age- base'
    ',
    t
    :l'
    I
    l,
    fri
    -3:
    I.
    -.'
    1t'
    : ::..:.
    lB,5

    li
    BUlOrt.,.F.
    ,1
    .
    and R.
    fuiderSon. l9/.1
    .
    tis.,,l,:,
    i),-r1
    ,,i
    lr;-tifi,i
    ,1
    lr:
    {
    jr.
    (,r,rr.,
    l::.,
    ,
    .,rr
    :,
    '.,liver
    parasites.
    Prog.
    i:ish
    eull.,
    4l
    l?i;;.i.
    l
    ilaqr-'10,
    R.. A:
    alt,j
    :l:
    A:-
    tranrl,.ri
    I li.
    1979,
    A
    cr:rrp.lris.,rn rif
    'p,rs1,i,1r3rri;t
    iq
    ,
    .tre1alode.(Sub:"omily:
    &reyrocephalin,re)
    poEirlar,ir:n't
    $t
    blrter;ill:i
    f
    ri_:,;i.
    ,'lieated
    and
    arnbieri!.
    areas
    of an
    !l
    iinois
    cooi
    i*9
    !,]ke,
    arirl
    ,1
    rrqr.r,'iry
    rrnh€.rtei1
    ,reser:voir.
    In
    fivalr;'rtion
    rt{
    a
    cooling
    lal:e fiiher),,
    vol,
    tit.
    !il.,iiat',
    llist,
    Survcy-Tinal
    rgpr;;.1
    to
    tha
    f
    leciric
    pcwer,?{sr:,lri;!r
    !nsf
    it_r"lle,
    pai,:
    Alto.
    CA.
    Hoffman,
    a
    striqeoid
    fi,
    L.
    1958.
    trcndtodc,
    Ixpltrirnr]ntal
    lostjq,Ji_gl5l,om,l1,
    studies
    on thc
    F.xp,
    r",]rfari,]
    p.)rasiL.,
    and
    i(
    met,lccr.ceri,r
    l
    }
    :?3-5.1.
    rf
    Itrlnter,
    G.
    W.
    ,
    I
    t l.
    1942.
    St
    rrcl
    ies
    orl Lire
    p{rrasites
    0f
    f
    reslr,vatrtr
    f isnot
    tf
    Cortnecricut.
    Bul
    l,
    Corrn.
    rreol.
    t{at.
    illst.
    Surv.,
    6l:I?B-BB.
    Lewis,.
    :,tlre
    rd..
    hody
    !4i
    anrl
    reight
    J.
    l{ickun,
    of the
    bluegil
    1g64.
    l.
    Ine
    pr.c!.
    ef
    fect
    irish.-I[Tr_;8fJ)-:fff_;-;
    of
    posthodipl0stonrun
    rnif}imrJrn
    rlprln
    "'
    sinrjefmanrt,
    c.
    ,J.
    1951.
    Parasites
    of
    f
    ishes
    of
    rrorth
    cDnIral
    i,l,]53,rclr,rli:Lt,s.
    I'lass.
    Dept.
    Cons.
    F isrr.
    Rept.
    ,
    ?B
    pp.
    S;nithermart,
    R.0.
    1963.
    Iffect of
    Lhe
    sLrigeirl Lrern,rl,rr.lu,
    posthorl
    iplostomrrryr
    rninimi{n,
    upon
    growtlt
    anrl mortal ity
    of
    biuegi
    I l,
    L*ponis
    rr"rcF'rocfil."f-.
    ,f,aA
    ,
    '
    EiTIilept.
    44, vol.
    5: lX/[-8,
    3S-0-398.
    i_
    |
    I
    ftiATi]ilil
    l"
    I
    1i
    i;
    T
    l,
    t
    I
    T.
    T
    t
    t
    t
    I
    I
    t
    I
    f'
    l,
    ::::
    t::
    r:'
    a:::.:i.'
    T
    tt
    rl
    .l
    fl.6

    pART
    I I tcoLoc
    r
    cAL I
    r,rvEsr
    r GAT
    I0Ns
    lor
    saonl
    cRt
    Ei(

    -
    :.t
    :a:::
    -...
    a:
    ,
    ':.
    ':.
    ;:.
    ,
    .
    ,.a..
    :
    :
    :.,
    j:
    :t:
    :'i:
    :
    :
    l'
    ..
    :
    ',
    'l:
    :
    :i::.
    ...
    Ltt'J I
    I !
    tt:.:'
    ,:
    ,ii.illiiti
    I
    ,
    '
    ,
    '
    IHI':ldfiiJCili)lr:
    ';iinh-L
    i.irt
    i-i:l
    .
    Ihe
    fasl
    Fork
    of
    thaal
    Crcer.
    I
    ies
    iumerJiatr:!y
    be]<:* Lrrir
    Cof
    f
    cerr
    i-.ri;r
    i.lan.
    lnil
    Strealn
    receiveS
    overf
    i0w
    f
    r0,n
    Ihrj I
    nke
    6J9r'irt!
    ilcr
    iod.) rlf
    nigh
    'ld.tLiJt'
    ,rrrr! irt
    t.ht
    pa5t'waterwaspUnpcdfrornthecreekinLrrtttr:.1akqtohaiLenlakefilliri9..
    0ne-year
    studi0s
    of
    r{nLL,r
    chcrnistry,
    phyLtp}.1riktqn,
    )ent.ltos,
    .lrrrl
    f rsir
    wrtr'.J
    i,nplenented
    io
    assess
    lhe
    ef
    f
    ect
    of
    'lake
    0ysrf
    :0rr
    .1qile,' pil,npinq
    0f
    d,ttcr
    f
    ''0n
    the
    East fork
    of
    Shoal
    Creek
    upoo
    t.its
    biola
    of tili-. sLredn.
    N[SCR
    ipi tt]t
    rtF
    lrlI
    sTt,i)y
    ARt_A
    ]
    The
    study,
    ared
    establ ishcrl
    0n the Easl
    Fork
    oi
    Shoal
    er-eek
    wal ,lirecLly
    soi,L,r'of
    Coffeen,
    Il linojs
    (T7N,
    Il3',,1,
    SecLiori
    23).
    ilith nn
    nv,3r'alle wi
    lLlr
    r'rf
    J5
    feeL
    .rrrrJ
    a
    gradient
    of
    4.5
    feut,
    rnile''
    -l',
    the
    cr'eek
    flows in'.t
    so,,+herly dir:c,"iori
    towai:tJs
    Shoal
    Creek
    and
    the Kaskaskia
    River
    (Lopjnor
    l9i0),
    ihe rlraindgr-.
    llrirnaril"y
    cortsists
    of
    pasture'or
    f
    drrnland whjch
    perir)dic,rlly
    ber:orncs inrirrtl
    iLeil
    .
    during
    periods
    of
    high
    discharge.
    Lopinot
    (1970)
    rr:port;cd
    thaL t,hr:
    sLr't?<rrn
    troLtorn
    consisted
    of 70
    percenL
    silL and
    30
    pcrcerrL
    sarrrl. HisLoricaliy,
    tirit
    stream
    provicled
    water
    for lake f
    i llling
    rlurinq
    a
    sever'e ,f,'oiqi,i
    wlriclr
    coincitled
    with tlid
    corqplq!ion
    of
    the
    Coffeen
    Lake
    Darn
    (CIPS
    1977l',
    Ifre
    auxilialy
    pt,irrpirr:.y
    "fi.iiitiei
    instai
    letl
    irnnrediate'ly
    belo',r Lhe
    Cofieen
    L.rkc
    Spi I
    lway
    were de.;ignerl
    to
    maintain a reasonably
    constant
    lakc
    levcl.
    Howcver,
    purnpin.r,
    operaLio'rs
    rverc
    apparently
    sporadic in
    the
    past
    and havc noL
    occurred at alI
    during
    the
    course
    of
    thjs
    investigation
    of
    Coffeen
    Lake.
    Prior
    Lo Lhe
    ortseL of Llris
    sL,rtly,
    trrr:
    last record of
    a
    .j
    spiliway discharge
    into the
    East
    Fork
    oi
    Shoal'Creek
    was
    ;lun:e
    r975
    (CrPS
    I
    s77).
    .:,::i
    ':I
    ::-t'I
    aa:
    '':,'=:
    5.1
    ,t
    ,t-
    :'
    -il
    il
    Figure l.l iliustrates
    sarnpling
    staLions
    Stat ions I
    ,
    2, and
    4
    were
    major .sarnpl ing
    piriphyton,
    benthos,
    ancl
    fish.
    Each
    of
    and a
    pool
    of
    the
    stream.
    Stations
    0,
    I
    for
    certain
    sutlprojects.
    on
    the
    [ast
    for'k
    of Sho,il
    Crer:k,
    stat
    ions
    f
    or
    wat.er^
    cheini stry,
    :'
    these
    staLions
    encornpassed'boLlt
    a'riff
    le
    .'
    r
    -
    ,'
    .
    b,
    ,lnd
    3 sr-,rved a5
    dncil
    lary
    staL
    jons
    Perry,
    and
    James
    ll.
    BuckIer
    I
    '.\:::
    1'5
    lAuthgr
    s
    :
    ":
    ,
    .'
    ,._
    j:t
    a:
    tt.
    t,
    I
    :;
    :
    .'
    .
    ':;
    ',1:;
    ';--'.t'.::t:';
    -::

    /
    ,/
    Lake
    ,/
    /
    //
    een
    4p
    Concrete
    Spillway
    Statigr.
    r!
    /
    'J,
    =Y
    Station
    2
    -:County
    Road 2OO0N
    q
    roo
    2oo
    ltl
    Meters
    Figure 1.1. Sarnpling, stat.ior)s or1 tlre
    llast t;olk
    oll SlroaI
    Cleek
    lrelos,
    the
    Coffeert l,akc
    rlam artrl spillway.
    :
    1.2

    :.a.'
    .,a:,,...:.:'l:=
    .'
    1,1,
    '::
    .
    ::.
    :.
    ,t
    t
    i
    t
    :'
    Stations
    2
    anrl
    3
    are ln
    close
    proxiftiity
    to onc
    dnnr.hrjr
    because
    J
    rubl)'l
    ,:
    rl
    Jrn
    w,ls
    onau,,,present
    between
    those
    two
    sites. llowever,
    in
    e*r-ly Fr:brrtrry
    l9i9
    t.h;r d*:n
    uas
    removed
    in
    preparation
    for
    the
    possible
    conslruction
    0f
    a
    Srnrrrarr*nt
    loa_hrad
    weir
    dam.
    Since then
    construction
    plans
    ltave
    been
    tieferre,J.
    Sl,ttir)rrs
    0,
    1.5,
    and"Z
    thus
    eonsilted
    of
    an
    tmpounded
    g001
    clur'ing
    nosl
    0f thc
    I
    year
    basel'ine
    study
    period
    on thls
    stream,
    while
    StaLion
    3
    reprcsent.ed
    the
    p00l
    jrlst
    belor thc
    rublile
    dam.
    :
    -
    .l
    ,.1
    t
    'l
    I

    :.:'
    :r
    ll
    '
    ttt
    '
    ,:r
    .,,1
    .
    '
    ,
    ,'.1
    Central
    lllinois
    Public
    Service
    Company. 197i,
    Thelaal
    dernonsLralion
    for'
    ','eoffeen
    Power
    Stations
    Units l'ra'nd
    2,
    Submitterj
    to
    Lhe
    !l
    iinois
    Pol
    lirl.ian
    .Control
    ,&oard,
    l,lay
    1977.
    ,'-
    Lopinot,A.G..l970..ibnt9o'[eryCountySurfacewdter:re5oilrces.Ii]irrois
    Department
    of
    Corrservalion, Springf
    ield,
    lll
    inois. I l9
    page.
    ,:"
    '"
    ,l;
    -t'
    ..
    "l
    "
    l,'
    'l
    'f
    I
    ''l
    'l
    ,,1
    "'
    .l
    :
    1.,
    ,I
    '
    l'
    T-.
    .
    :'
    "
    ,
    .
    l't
    -':r.
    l=
    "rt
    t
    ,l
    ,':
    'l'
    .',
    '
    1.4

    stcil0N
    2
    SIIOAL
    CRIEK I{AliB
    QU'\LJIY
    h,,,'J
    ,,Sarall
    Liehr-5torck
    :
    fiBSTRACT
    An
    jnvestigation
    of the
    East Fork
    of Shoal
    Creek
    was
    conducted frorn
    August
    lg/S
    throlllr
    septernber'
    1979
    for
    the
    purpose
    of
    srutlying
    the irnpact
    of
    Coffeen
    Lake
    ove.rf
    lo:t/
    0n
    Shoal-
    Creek
    water
    quality.
    In
    the
    f
    al
    I
    of lgZB,
    ilre
    sarnpi irrg
    station
    upstreatit
    of
    the
    Coffeen
    Lake discharEe
    h,rd
    very
    hiEh
    conccrrLrations
    of
    clrloride,
    total
    alkal inity,
    soluble
    orthophosphate,
    dnrt
    0rg.rnic
    cer5o1
    a1d
    iow
    concentration
    of dissolved
    oxygert
    relative
    to
    l.he
    other
    sarnplinq
    sLrrtions.
    These
    concentrations
    indica'ue<J
    that
    Lhe
    streanr
    was irnpacLed
    by
    dn
    upstre,drn
    sewaqe
    effluent
    durjnq
    that
    periocj
    of low
    f
    low.
    In
    ln
    ,1llt
    tl(lt-cn
    ]9/9-Llt-e_
    ty
    /y,
    tne_.--tp-Llllg
    sp-11g-f
    lgocl
    na0
    had
    .---:_*__#
    more
    more
    effect
    etfect
    on-_![q_!a!er
    on
    the
    water
    quality
    ty thtthan
    did
    the
    overflor:frorn
    Coffeen
    Lakr:
    CharacterisLics
    of
    Shoal
    Creek
    water
    qual
    ity
    at Lhat
    t i,ile rvsr.{,,
    0xtrernely niqir
    turbidity,
    along
    with
    high
    levers
    of iron,
    phosphorus,
    dnd inr.lrganic
    rritr-ogerr,
    and
    relatively
    low
    concentrations
    of
    chloritlc,
    alkaliniLy,
    harrlness,
    arrrl
    sulfate.
    By
    April
    1979,
    Coffeen
    Lake
    overflow
    alterr:d
    iidter
    ctu,rlitv
    at Shc,
    cal
    Q19_ek.
    The
    ne rno_siJ9l_'cedb-te
    most noticeable
    ef
    ef
    !'ec_tLi_gl-e_i|Iqq_t_e_f'
    fects
    were
    rtreater
    S_qjl-l_!e
    sulf atp c
    c_orrcenLrat iorrs
    ..rrrrJ
    lower
    '"::"'
    glll-l-inilf-ut
    !n9-!9ry!:l$gq*:!guqlu.
    L"s r/vw'rJer
    sqrr
    lisll-vr:.
    ny-.d}lrlli
    Bv-.Jpng..![o-!qJf-f,eqts,ve,',]
    Llose
    ellects ivgrg
    no
    n0
    . .
    *.*-*_-.-:
    l.919gl-l-Yident.
    The
    summer
    f lood in
    August
    l9i9
    harJ
    a
    s
    tight
    rti luting
    ef
    f
    e
    pat
    orl
    Shoal
    Creek
    water
    qual
    ily,
    but
    Lhere
    were
    only
    rnirror
    intJicalions
    of
    irnpac,u
    by
    I ake
    overflow.

    ,ilIt{00uc
    I
    l0ri
    Thls
    investigation,
    con<trlcted
    bet-,veen Aur;rlst
    lllii
    r:iri',:]g;ir,.':,i.,r'r
    il,r,.r,
    ,rr:1.:rr;:j?i
    with the fol
    lowing
    cbjectiveg;
    ,'
    bo determine
    ili
    ths
    ialart
    .rr
    i-ir!
    f
    ,-,rrr
    l*.e
    Overf lon
    Orl Shoal
    Cree k watcr
    qual
    i
    ty,
    (?
    )
    ,.n,:
    rnp,1i'l
    r.t,(.
    g\tai)i
    r:.;
    w.i,1
    1;r.
    I
    ri;c
    Shoal
    Creek orr
    the
    rtoter
    quality
    of the
    lake,
    dnd
    {3-!
    lnr-.
    rnrplr-!
    ,if
    i.iri:
    ;ri'-,iin,;
    operation:on
    thr' water
    qual
    ity,gf
    5hoal
    Crr:ek. 5'i nce
    witqr
    ,{d,,
    rr,l,t
    r.1
    J.rprj,.i
    :iit:j
    the
    lake
    fron
    Shoal
    Creek during
    the
    sturly
    per
    ic{J,
    only
    liie
    iinp,lct
    ,lf
    t-:rrt
    lli.il
    over'f low
    on the
    sIrearn was invesLigatert.
    Tlr
    is
    -lb3,ict
    ivr,
    n,rs
    ,]-i. ]!i!1
    i
    r irr,':l
    it-r,
    dets:rtnining
    the relative
    impacts
    of
    ldke
    ,tater
    anrt
    il
    irnat,:1,1r1 i,'-
    rl
    r-.v.:rir,tr
    (,r-ii
    Lc
    ternperaturre,
    f
    loods,
    arrd
    'l
    ow
    flow
    on
    water
    qualil.;,
    pJr,:rrittqr'!
    i:r
    ':,n,rJi
    ir,,,l;:,
    a:,-a
    -a:
    :
    -=
    l=
    l,'
    ,.
    I:.':
    I,"
    :..:.
    l'
    T.
    l,t
    .
    l'
    t
    I
    T
    l'
    t
    T
    :
    l'
    2,2

    ,
    '.
    ,:'
    lAllPl-
    t
    r,1f,
    !{-rqi
    ir
    j,
    i
    tldLt:r
    QLI+l
    i
    Ly
    ?ar,Jni-'tr.rE,
    *r,.,{
    ,'
    !,...i-.
    it
    .-
    I ::
    s#,Jttf
    qf
    :lir,i{ifi,rj
    ii'!.
    i.l;.
    ..:,
    i',.;;r,r='t,
    2 al{
    3
    t4',
    .t!',tv)vr:
    J
    ,1i,1
    t.iu:r..-1
    .i{..,;r
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    :..;..
    \ti'
    :,:.
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    rt{.it't|
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    Lhlliltttpnat€r
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    :l:.i.,
    ;
    ;i,..r
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    rlrld5ilrL.rl
    twir:ri
    g4qlr
    ,nOntn
    dt.in;
    t?r,;
    St:l
    Jll
    l)+ri,,il.
    t
    ILLi)
    rr'1il
    r\llirLYT
    lCAt.
    14t
    1,.t0il5
    l'later
    S6nples
    iler€ col
    l€ct.rt(l
    in
    pl,rst
    ic i;ot.Lles
    aptrrlr
    i;rlc! r-,
    1y
    :1
    .t,
    111;11,';-.,i
    itr:'l
    i.;a
    thLr
    SurF,tce,
    :)amples
    anilyzctl f
    or
    Sol,tltle
    ir^r)rl
    ^errl
    i,nrrr-,rj
    i
    al
    ,tI,J
    f
    i
    itt,.ii,i
    arrr!
    fixerj'rriLh
    hydruchloric
    acir!,
    and
    garpl,lg
    for ferrorla
    irrrr
    vrer-e
    fir.€{l
    irwn-erliately
    with hyclr'ochloric
    acirJ
    ,
    Saitiples
    an,liyzed
    for
    sulf
    irlt:
    fur
    l5
    ..Jere
    .
    f
    ixerJ
    irn:nedintely
    witlr
    0 ,2
    N
    zi
    rrc
    acettti.e,
    .trtrl
    analysts
    yJ(?re
    r,.lrr
    wrtirirt
    l4
    lroLrrs
    of colle:Lion'.
    Sulf
    ite
    ,anr!
    0H
    vJ{lre
    ilqa$u
    retl
    .itr
    tlie
    f ieirJ
    ,rr
    in l.rtr:
    labrtratori,
    lrnnerliately
    af ter
    collectiorr.
    IenperaLrrre
    ,tnd
    tJissulveJ
    oxySen ?rrlr'rr
    ;rrrjJ\i,r.,.,,1
    in
    tlte
    firllcl
    with
    a YSI
    l4odel57
    dissolvcrJ
    oxyger'r
    rnrtLcr'.
    (,orr<luctivity
    w,ls
    rnij.r.,urr,:l
    in
    the fielrl
    wiLlt
    a YSI Moclel
    33
    S-C-I
    nreter,
    anrl
    Ltre I'r-,sulLs
    w(:r.rl
    c()r.recf
    ,j(l
    f,)r
    ternperabure dnd
    used
    bo calculaIe
    toLdl:dissolvecl
    soliris.
    Chernical,p4r-4r1s1g1.5
    'de'r-e
    tnerlsrtrecl'.
    using
    the
    .sarne
    analyf ical
    rnethods
    qiven
    in,Secl;
    ion
    3,
    pa,-L'
    I
    ,
    .
    her-ei
    n,
    -
    2.3

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    iii
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    I SSOLVEO
    SCL
    II)S
    .
    :
    '
    l
    -
    l
    Iotal
    dissolvetl
    s()lids
    (1DS)
    concenLraLir)ns
    ,verd rnrrctr
    hi,;trr:r
    nt
    ,it
    .rLi4n
    i,
    ,
    t'
    upsLrearn
    of
    l,htl
    lake
    discltarge
    channel,
    in Lhe
    f
    all
    o
    r
    l!/\
    Llr,rrr
    at
    t.trr-,
    oLlrCr.
    sbaLions (Fig.
    2,5), TDll
    levels
    increasr-,d
    Lhr.rrqrr
    Lirr: vri'r.cr-
    (rt
    nrr
    .,r,,,
    ,,,,,r,
    then .Jecreased
    subsLanLially
    in
    l,larch
    1979
    wiilr
    Lirc
    5pr'inq
    f
    lortl
    (Talrlr
    ;:.?),
    In Apri
    1,
    T0S
    levels,rose
    at::all
    staLions,
    wittr Lhe
    gfeoLesL
    irtcrr.,a5rj'i)(:crrrrirr,:l
    af
    ,italions''idownstream
    of the,Cof
    fee'n
    l-ake
    sp:i llway.
    AL
    thaL
    Lirrrr:,
    l'i]S
    t'
    concentrattons
    werc
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    at:
    jtaL
    ion
    0,':l.tre
    I
    akc
    over-f
    lrl/,
    clld.r.tf
    I
    ,
    lharr
    at
    the
    Shoal
    Creek
    ,stations
    s
    jnce
    If)S
    was
    typical
    ly
    gr.g,lt-Jr.
    in
    Cof fcen'Lake
    Lharr
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    z,=
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    J.
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    :.]
    i
    n,
    Shoa'l
    Cr'eek.
    ',Ey
    rn!d-l4ay,'
    however,
    .the
    c0rr(.erllr
    ttniforrrr
    dtnonq
    staliL.ns.
    Then in.l.'rfrt,r]uly
    ail(l
    enr
    attrJ
    ovgrf
    lor f rorn',thc
    I
    ake occurred,
    T0S
    levoi
    s
    i
    n
    i
    substanLiatly
    while
    lr:vels
    at St.ation
    0
    irrcr.cas.lC.
    rtat
    ion, lavels
    of
    T0S
    .Jere
    back
    to nor-,na
    l,
    rvil.h
    rro
    overflovr.
    Sul fate
    concentrat ions
    Lake. In
    Shoal
    Creek,
    Stations3and4frorn
    alions
    hat,
    aqain becrrnc f.lirlr'
    ly
    liurSrrst-,
    whi!!t
    rtrl0t"itrr'
    i
    :.)riil
    Shoal
    Creek aq,:in
    .itari:i.-ir:iJ
    lly
    nrirl-Augrrst.,
    llrp,lr.i
    r
    '
    ,:
    app,lrent.
    el
    f
    r-rCt
    f
    r0:n
    tric
    I
    li:*
    Alkal
    inity
    concentratlons
    were
    cortsiderabiy
    gru.atelin
    Shoal
    Creek Liran
    in
    Cof fcen'Lake
    at al
    I tinres
    except March
    l97g,'durlng
    the
    spr-in3
    f 1oo11. Prior
    Lc,.
    M.irch
    l97i),
    alkal
    inity
    was
    observerl in
    the
    rangc
    ol' aboirL
    250
    tri
    2il0
    ;,r3/l
    .rs
    CaC03
    at
    S'.ations
    3
    and
    4, however, frorn
    li,ugrrst throrrqlr
    l,love.nbcr'lg7i.l,
    lr:vels
    *ere
    3C
    to
    90
    :ng/l
    higher
    at Statiort l.
    Thcse rlaLa
    c0rrcspond to
    tri3h
    Lr;tal
    dissolved
    r;illids'levels
    at Station
    I
    cJuring
    ilrat Linre;
    ln irnril lst/9,
    alkal irtity
    i
    ttcreased
    at
    al
    I staLions
    in
    spite
    rlF
    over'f
    lol
    f
    r-o;n
    trre Lrke, buL ilr,l
    levels
    at
    stations
    rJownstream
    of
    the spil lwa.y increasr:d
    Jt ,t
    slower'
    r'dLe
    tharr
    dirt
    concerttratiorts
    at
    Station
    l.
    Alkalinity
    levels
    continrrcrJ t()
    incr.r],tse
    unL: l
    July,
    and
    a
    small
    dco-easc
    was
    ollservctJ
    after
    the
    sunrrner.
    l-lootl
    (table
    2,2).
    ilar<Jness
    inct'eased
    at all
    stations
    rlurinq
    the f
    all of
    I978.
    As
    rvit.h oLlrrr
    cl issolvetl
    contpr)rtents,
    concerrtrations
    decreaserJ
    rlranrrt.ical
    ly irr i,1ai'cir
    1979
    drrrirrq
    the
    sprinq f
    lood, then
    started increasing
    to rtorflral icvrtls
    in
    April
    (Tolrle
    2.2).
    Ch)qr
    ide
    cortcentrat.ions in
    Shoal
    Creek
    erere
    gcr'teral
    ly
    si:nil ar
    to
    concenlraL
    ions
    i'n
    Cof faen
    Lake; therefore
    spi
    l lway
    discharges had
    I
    ittlc
    ef
    f
    ect
    on
    the
    sLr-crlrn.
    Al
    I
    stations
    ,lxcept
    Station'
    I
    had sirni
    lar chlorjrle
    levels
    (Iable
    2.
    |
    ).
    Chlorirje
    concentrations
    at
    Station I were
    extrernely irigh rclative
    Lo the
    oLlrer
    sIations
    frorn tlte beginning
    of, the study in
    August
    1978 tnrorrqh
    l'lovernbcr
    l9/3,.,uiLn
    Iirc
    rnixjrnum
    concentration
    if
    145 rng/l
    observeri in
    Septernber.
    Station
    concerrtraLioni
    trctP fairly
    cortsist.ent through
    tne
    winter,
    then
    clroppert
    Lo
    rr
    minirnurn
    of alrorrl lZ
    rlrg/l at al I stat
    jons
    in
    llarch 1979
    (Table
    2.2).
    Frorn
    thaL
    time
    urrti I Llrc
    enil
    of
    the
    sturly
    in Septernbei 1979, Station
    I again
    harl
    the
    h ighcst
    chloritte
    concentrations
    with a
    rnaximurn
    observed
    in
    Septernber
    of atlout
    55
    rng/1.
    -
    I
    I
    I
    l
    I
    I
    r
    r
    I
    T
    t.
    !:
    I
    I
    1
    t.
    l'
    !.
    ':
    -::
    !.
    :.1
    T'
    I.
    t:
    l.
    !,
    :
    I.
    f:
    :
    l.
    I,
    .
    l.
    l,
    ...
    l:
    l.
    l:
    ''.'t:
    i'
    .
    :::,
    ..
    !:
    l.
    in
    Shoal Creeki'were,,lconsi'cleratrly
    lower
    than
    in
    Coffeerr
    ccncentrations
    were-'slightly lriglrer.,rt
    StaLion
    I t.hat
    at
    August thiough
    Novernljer l97tJ.
    :Frorn
    Decernbcr
    to i,4ariir,
    2.16

    c0ncentration5
    at
    al
    I
    threc
    slrca,l
    stationi
    ,rr{.r^il
    fii:el i.y
    tirr:
    ia:re,
    i
    {lrt(:r
    rdse-
    :
    vlaS
    Observt'd
    in
    llarChr
    C0r'rqr5pgnrling
    LO
    tlir:
    .,;rr
    iri.l f
    ]i-ri;1j
    i l,:5 i,:
    1.
    ?
    j
    .
    lri
    i1pl.,i
    j
    drld
    May'
    concentrdtions
    *erc
    rnrrch
    lriqher
    aL SLdtirrn\
    3
    anrJ
    ,i
    tii,lrr
    ii
    .:r r..;in
    I
    ds
    water'
    from
    coffeen
    Lake harj !'llore
    irtfluence
    on
    sLre.Jr:;
    c*rrf*n!.r,lt.i*n.,.
    siili.rte
    levels
    were again'similar
    at
    all
    sLations
    friin.Jrrrr,:1979
    to
    the
    cnc
    nf the
    sLitcy
    'in
    Septe;nUerllg/9
    in
    spite
    of lhe
    lake
    ovtr'f
    lo*'i
    nq.durir'lij
    th,-.
    iii,iiry1r:r
    rrgo{j
    ':
    I
    I
    I
    l-
    ::
    'l
    :.i
    'i
    :1,
    Thesc
    data
    for
    the
    cornponents
    of dissolverj
    solids sugjesL
    t)rat
    cspecially,rrjversely
    inrpacterl
    by
    the
    loir
    flow in
    shoal
    creck
    i
    fall
    of
    1978.
    The
    high
    alkal
    inity
    anrr chroriic
    levels,
    alr:nq
    organic
    carbon levels
    and lor
    dissolved
    oxyqen,
    irrdicate
    thal
    upstrearn
    of
    Stat
    ion
    I
    probabl.y
    caused, Ure riegrarlal
    jon
    of
    .rra,r.er
    P
    I{OSP!IORUS
    SLaii':,ri
    I was
    rt
    l.he
    s
    ilnnre
    r'
    .
    .t
    n,.l
    ,ri
    t.h llre
    lr
    i
    gher
    3
    SCwrlte
    4f
    t
    luefli
    ,
    qual
    i
    ty.
    'l
    I
    Soluble
    orthophosphale
    concenLrations
    ,,vei:c'.vCr."v.
    hi3h
    aL
    a!l'-sfrLicns
    in
    ikr-,tr,rrbcr:
    1978,
    brrt
    especial'ly
    at
    sLatiorrs
    I and
    3
    (0.
    .t2larrc
    0.27 nra ?/t, .espel,t";;;i,
    Concentrations
    increaserl
    at
    all
    stations
    in,4ar.cir
    l97g
    to
    aborrt.0,l5,ng
    p/l
    (Table
    2.2). Total
    phosphorus
    also
    irrcreased
    at
    LhaL
    Linrc
    (rnca,r;
    0,64
    i,rg
    p/l),
    corresponding
    to
    extremely
    high tur:trirlity.
    l-otal
    .phcslthorus
    levr:ls
    i'lg.rdrr)d
    after tl'trl
    stltnmer f
    lood in
    August
    1979
    (rnearr;
    0.59
    rng
    P/l),
    lrLrL
    orLtrophosph.rLe
    levels
    remained
    low
    (Table
    2.2).
    The'
    phosphorus
    concr:rrLration
    was
    i,sr/il
    ly less
    than
    0.1
    0 mg P/
    1
    .
    {
    I'ROGEN
    ,
    Concentrations
    of inorganic
    nitror;en
    weru sirnilar
    diron0
    sLre,rnr
    sLations
    aL ail
    titnes:
    (Table
    2.1).
    'Anmonia
    levels
    were greatest
    in
    Janrrary,
    February,
    i,,ri
    i4ar.cn
    '1979'anrj
    least
    during
    the
    suqlner'
    an,l f ati
    or
    l97tl
    {tante
    Z.e).
    Co,r.*uriuiion.-
    were sliqntly
    increased
    in
    August
    lg79
    after
    lhe
    surnner
    floorl.
    !
    Concentr-rrLiorrr
    wrJrtLg
    of:
    nitrate
    were
    low in the
    sunrrners
    anrl
    in
    Lhe f
    al
    I
    (O.l
    tc
    0.6 nrg
    tt/1,i,
    hut
    '
    increasecl
    dur:ing the
    winter.
    Maximurn
    concenLra:Lion's
    werc
    ol)sr'rverJ
    in'Mar-cir
    an0l
    npiit
    (Tatile
    2,2).
    A slight
    increase
    in
    nitraLe
    occurrerf
    at
    Stril,rn*'3:ari,l
    4],
    i,r
    Augus:t
    J979
    af Ler the
    su'imer
    f
    loorl; apparently'causerl
    io'par^trby.ovorr.lil',,j
    f ,.o,rr,tl
    cof
    feen
    Lake.
    Ni
    trite
    concentrations
    rvere
    low
    (
    .0r
    to
    .0.t
    nr,J
    i{/l
    )
    at
    ,rl
    I
    t il,res
    .::
    1,,
    :I
    ;.
    -
    i.':tl
    -
    ,=:'
    :.:]
    .:r:rt
    ;:;:i
    ;::i.I
    2,17

    a:
    tr1
    F,l,
    Sir:.='
    F;t;
    .
    g-Yr:';
    .'-
    :
    fr.
    $
    f;i''
    F'1i....,
    El-._ .
    :
    :rl:
    '
    ::
    :
    fi..a
    a;':
    ,
    f.'::
    .'.,:
    i1:=:,
    .,
    ,:,.;,
    l:,t.,t
    '
    'i;;'
    ,
    '''
    al;:
    ,,..:
    .
    {,;,',"
    L.:..','
    ?;:'','
    '
    :r:' -.'
    i,,..,',',
    ,
    '
    ii:
    ,
    r.,:'
    ia,.
    r'.
    'i:
    :'
    :',
    fu
    t''
    .;i:"-,
    it"
    i;ij::
    *.:::..
    :;:-:]:.
    :'
    !1'..."
    ,
    :;,:'.:'.
    "irt
    t";','
    -
    i:-.:
    t,'-
    a
    {7.'.,
    ;ii"t
    'j
    it,,i:
    r
    '
    ...
    '.
    ':..
    iL.,,-,'''
    ;i:,;:.:
    ::
    iti:
    1
    !i,t',"
    '
    '
    i!,t,-''t
    '
    "'
    i;;'.:,..
    :
    Ihe
    overall
    impact
    of
    Coffeen
    Lakc
    overflo* is
    difficulf
    to dtsess
    sincc
    '9ver!!ryyo.gY.r':gg*s:.:-tg-.p-grts=!s-9l].rlg.!-ducjlar:ge,,n.,nour.|
    Creek.
    a-_
    Parafeters
    that'were
    found in
    especial
    Iy
    irigh'or
    loy,
    concenLr,atiorrs
    in
    =%
    except
    during
    ilarch'and,Apri
    l when
    levels
    .rere
    0.09
    and
    0,04 to
    .05 mg
    ti1l,
    respect
    ively. Tire
    total
    inorgan
    ic
    n
    ii,roge
    n
    concentrat ir:n
    was greatest.
    1n
    ',!1r
    ci)
    1979
    (2.97
    ng
    N/l) and
    lowest in
    Augusr
    l97B
    (0.09
    mg
    r{i
    l),
    !{
    f_ej!,f.eleJelsr
    tue
    -rq@J-ree!
    {
    t
    o
    t'
    l- ;-;i;;;
    -;"t,
    d
    i,-
    i rl i i r.,
    u n J
    --
    alkal
    inity)
    had
    limited
    impac!.3lLt.he
    receiving
    stream because
    of
    the
    high
    discharge.
    station
    to
    --
    lake
    --'-
    .''
    was
    overflow
    Changes
    very
    were
    sm-Oll.
    in
    of
    the
    During
    short
    levels
    rJuration.
    most
    of
    of
    these
    thc
    Temperature
    paranreters
    sLudy
    periorl
    at
    increase-d!-.the
    rJownstredrn
    al l
    stream
    stat
    st,ations
    do,wnstream
    iorrs
    lrad
    tlue
    -G-#
    s irni
    I
    ar
    concentrat
    ions
    of most
    parameters,
    or
    t.he-
    wo_ts-l*$!l-f_g-ual
    ity
    was
    f
    ound
    _
    _y
    at the
    upstream
    stat
    ion
    -
    -*
    --:*'*-***"-
    /
    \
    2.r8
    :
    .:
    -l'
    :.::
    I
    'l
    'l
    :
    'l
    :'
    I
    f,
    'l'
    'l;
    t
    I
    I
    ta
    . '.'
    ,
    :.a .
    tl

    '
    L
    l
    I[RfrIi]Rt
    ,C lI[{)
    American
    -'
    '
    Sfandard
    Public
    i{etlrods
    Health.Assgiia!ign,
    for
    the
    E.rarninatlon
    Arnerican
    of
    ware,'
    Hater
    ur,r
    torks
    xriiuruinr,
    i'rssrltiatirln.
    lgl6
    -iir"
    ..,
    ..
    tlashington,
    0.C,.
    B74
    pp.
    :
    ,
    ,
    :
    '-.'.,t.
    ,
    ,..:,1
    Hdrvel''ll.'li.
    1960,.
    'Ihe
    chemistry'anrl::flerti-l
    :.
    ity
    of sed
    w3ter5.
    Carnnrrtlge
    Univ.Press,London.24app-,.ii
    Jirka,
    A.
    M',
    and
    li.
    J.
    carter.
    r9i5.
    Anar.
    che,n.
    4z(g):r39i.
    '
    ".
    Park,
    P.
    K.
    1969.
    Oceanic
    c02
    system:
    An
    evaluation
    of Len
    rnethorts
    ot
    .investigation.Lirnno.l.0ieano9r.la:l79.
    |
    U;5.
    | a
    G{i;iogical
    h.ia-!
    SYrvgy.
    l978-1979.
    l,'latcr
    Resources
    Data for
    Illinr:is. 14at.er
    Data
    Report
    IL-78-1,
    and
    IL-79-1.
    .
    .:::
    r:i.::
    2.,1
    g

    PTRIPHYTOH
    O}I
    stcr
    10il
    3
    ARIIFICIAL
    SUBSIRAT[5
    IN 5}IOAI".
    CiigII(I
    by
    Larry.
    ll.
    lCout,anl.
    ABSTRACT
    Perlphyton
    col0nizing
    artificial
    suhstrates
    exposed
    quarterly
    for
    ZB
    clay-periods
    from
    August
    l97B through
    May 1979 were
    exarnined
    frorn
    four locations
    in Shoal
    creek
    to
    determine
    effects
    of overflow
    from
    Coffeen
    Lake,
    a
    cooling lake for
    a
    coal-fired
    power
    plant.
    fire
    relative
    importance
    of
    the
    major
    divisions
    of
    olgae
    was
    similat between
    most stations
    on
    rnost
    sampling
    dates
    but
    the
    dorninant
    division
    was different
    between
    col
    lections.
    The
    greatest
    diversities
    of a'lgae
    occurred
    during
    August 1978
    and the
    lowest
    ones occurretl
    during
    Novernber
    lgiB;
    Station
    4
    generally
    had
    the
    highest
    diversity
    cluring
    the
    quarterly
    oeriorJs
    when
    samples
    frcm
    that
    station
    were
    collected
    .orfi.q
    th-
    rerird
    !L.did
    .999-U|*ff9a_-Cofleen-
    Lrke,-in-Apri
    l-and-!rlay-*1,97.9,-!he*peciphy1.qt--9_olllry!.i!L
    ::,i'
    ,':,,:.,,.
    :f'
    -r
    li-...
    itl'
    .t ,
    i'1,
    downstream
    frorn
    the confluence.of
    the
    over'lilow
    frorn
    Coffeen
    Lake with
    Shoal
    g-ly:@,j:*_gl_"rg!,
    !r
    tnl_:r!ru*.
    ;1 ;i{;;
    i,;;}il;;
    i;i;.
    3,1

    irr,
    -t
    F+
    :.r:
    .
    i9.;'.
    '.
    Fi''r: i,
    l=,,'
    *1:::
    '
    $=.i.,.
    ,',
    Ft!:j.,
    _
    -
    :,
    F;::.
    :
    i+r,,r.
    .:
    i,t,''
    .
    I'j
    ii:::'
    .
    :
    ii::,.
    '
    i::'.
    '
    +,',
    ;t.r
    ":
    i:i,
    ,
    '-,
    :
    ::l
    ii'.
    '
    r:.
    ii: ;
    i:'.,:
    ':-,::-
    ,
    .;a:
    .
    :
    ;:a
    .
    i:,.,'
    :s..-
    .
    litr.
    ,
    7;,'
    t':
    :;:
    ti
    'ii
    t'
    ,
    i::1
    '.
    1.j
    :
    ,i:i.
    i::
    :;j:
    ;:.,
    ,
    .,,
    ;,:"'
    '
    ;.,'.
    ;j,j,
    *r.,
    ,,:,
    '
    t::
    :
    =:.
    i.i.
    '{;;'.:.
    :
    ;
    !;:'
    L.:.
    ,
    {,t';.'
    i.,'
    ,
    '!a::
    ,t,,:
    -'.
    ?:_):
    {:;
    ii'",
    -:.t:
    .
    .:.a:
    ,:!
    ,
    :a:"
    ::
    ::
    il,,:
    T
    T
    I
    T
    I
    t
    t
    IIITROI)UCI ION
    Attached
    al0ae,
    or
    periphyton,
    are
    an
    excellent
    means
    of aseessing impacts
    oi'
    point
    sources
    of'
    pollution
    ort aquatic
    environrnents.
    Because
    0f
    their'sessile
    natuFe, the
    algae
    dre
    confined to
    a
    specific
    area
    such that the
    algal community
    th'at
    develops
    is,ind'icative
    of
    the
    environEental
    'aonditions
    in
    that area. To
    fuither
    l'imit
    the
    number of varlables,
    artificial
    substrates such as
    glass
    slides
    can be used
    anrl
    placed
    in
    suCh a
    position
    in
    the water
    that
    variations
    in
    sLtbstrate composition
    and
    I
    ight
    penetration
    betr+een
    locations can be minirni zed.
    In this
    study, Shoal
    Creek
    periphyton
    colonizing
    ari
    ific'ial
    substrates were
    examined
    to
    deter:mine the
    impacts
    of
    rverflow
    frorn
    Coffeen Lake
    on Shoal
    Creek
    unO to
    determine
    over'al I
    dynan,lcs
    of
    ,the
    Shoal
    Creek
    periphyton
    comrnunit.y.
    Salnpling.stations
    v{ere located upstream
    and downstrearn
    fporn
    tlrat
    point
    at whiCh
    water
    frorn
    Coffeen Lake entered
    Shoal Creek
    to
    all'ow
    cornparisons
    of
    unaffected
    and
    potentially
    affected
    areas.
    -
    MATERIALS
    AND HTTI.|ODS
    Periphyton
    was collected
    quarterly
    on
    glass
    slides suspended
    in
    floating
    periphyton
    sanplers at
    a
    depth
    of approxirnately
    0.025
    n
    for
    28 clays.
    ,Quarterly
    col'lections
    were made'in August
    and'November"
    l97ii;
    and February
    anr! May
    1979.
    Two
    stations were
    selected
    on Shoal"Creek;'StatioR
    .|.5
    was
    upstream fr'orn
    the'
    ,'
    ,l:
    .
    .
    .
    a
    junction
    of
    Shoa'l Cre-ek with'the
    pumphouse
    canal
    -,
    and Station
    2
    was
    locaterl
    below
    that
    junction
    (see
    Fig.
    l.l, Part
    I'l). THo'addjLional
    statiorrs'were
    chosen
    for
    study frorn
    l,lovernber
    1978 through May
    1979;-station
    I was
    300
    m'upstrearn
    f
    rorn
    Station'1.5
    antl Station 4
    was
    located appr-oximately
    one'rn'ile
    downstrearn frorn',
    :
    Station
    2. At
    the tirne
    of
    collection,
    each of three
    sl i,Jes was
    preserved
    in'[act
    ih,a bottle
    containing
    2 ml of acidif ied'Lugol
    !s solution anrl
    63
    rnl
    of,water.'
    .:
    l'
    t'
    t
    T:
    l'
    I
    T
    -l-
    :l'
    ,:,
    ,t,,;
    ''
    ,'':,
    ,..
    1-
    I:It
    .
    ,
    ::::
    :i
    ''
    i=
    tr,'ii

    Scraplngs
    f ronr
    an
    entire
    sl
    ide
    wr'r'c sLirri:ii
    irr
    a
    i,iarirro
    lilenr|:r'
    drrd
    rlilir',-cd
    to
    a
    known
    vo'lume
    (e.!.,
    50
    ml
    ).
    Duplicate
    sainBles vlcre
    then couni.ed
    by
    thi: same
    procgdure
    outllned
    for
    plrytoplani<fon
    sarnples
    (Secr"lon'5,
    part
    ll,
    r',c,.oiri1,':g*rrt,
    that the
    appropriate
    rnultipl
    icatlorr
    f
    acLors
    .*cre
    used t.l
    qivr:
    densiLies
    'in
    alqal
    units
    per
    l0
    cmZ.
    RTSULTS
    ANt)
    DISCU.SSIOI,I
    c0r'rPos
    I T
    I0N
    Sev-e11ty-,fi.v9---a-ig.E.l--taxa
    were identified
    from
    collections
    of
    peripnytcn
    from
    artif
    icial
    substrates i:n
    Shoal
    C.reek
    (Table
    3.1). As
    expected,
    .nany
    of
    those
    .
    same
    taxa
    were
    found in the
    Coffeen
    Lakc
    periphyton
    collections
    (Section
    5,
    par.t
    l,
    nereln).
    ABUNOANCE
    Sinilar
    periphyton
    dens
    jties
    were
    present
    .at
    Stations,
    .l.5
    and
    ?
    <Jurirrq
    LuqusL
    l97S
    (Table
    3.2),
    The
    periphyton
    densities
    decl inecl
    frorn
    roughly
    2,5C0,000
    units
    per
    10 crnZ
    jn
    August
    to
    appl'oxirnately
    1,200,000 by ttovembe,
    197S
    (Table
    3,2)' Stations'l
    and
    4 hacl
    algal
    rlensities
    of
    approximatel.y
    234,000 and 869,000,
    qe'spectively,
    jn
    Febf uar
    y
    1979; tlrose
    densiL
    jes
    were
    'the
    lowest..r'ecordecl
    f
    or
    the
    .*.;--;,--'-.
    study
    year
    and
    !.9"re_U9_s_L.-!]-fgly*a__resu.l_t_o_[_teduce_d-],1-qht
    avai,tiibit
    ity
    and
    .
    lergg.atgfg_ttd-qr_99qlli-LUF"d.!_o;lgygt
    9f9W.!! rqles.
    The dcnsiries
    of
    periphyton
    in
    l'1ay 1979
    were substantially
    greater
    than those
    recorclerl
    in
    November
    lgTB and
    February
    1979
    at
    alI
    statjons;
    Station
    I
    had
    clensities.of
    2t326,200
    and:Stations
    I.5,
    2, and 4 had densities
    0f,3,038,510,
    4,744,40C;
    and
    6,'148r360,
    respectively.
    '.,'
    ::
    1,:'
    .
    :.',
    j..:-
    '.t''
    The,-r:elative'
    importance
    of
    the
    major
    stations on
    any
    given
    sampling
    date,
    different
    in nearly
    every
    collection
    dominant
    group
    in the
    August
    l97B and
    "=;',,,,:.
    l:.:
    -
    :
    '
    .:-.:::
    :
    ::-
    ,
    .,,,a
    ,
    ,:,
    ..
    .
    '
    ',:
    .:.'
    ..
    ,
    '1,
    .
    : '.'
    divisions
    of
    algae
    was
    simi lar betweert,no(t
    but the
    most
    abundant a'lgal div'ision
    was
    (Table
    3.3) . The Bac
    il l
    ar,iophyta
    were
    t,he
    May
    i979
    collectiorrs, the Chlorophytd
    t.rere
    \
    -
    .1.
    1a

    '-'
    Iablc
    3.1.
    Algal
    t,rxa
    identifierl
    from
    quarLerly
    periphylon
    ,
    col lections
    f
    rsin artif icial
    srrhstraies
    arrfour sanpi
    iriri
    stations
    in
    Shoal
    Creck from
    Auq,rrt
    lgZB.ihrcrlrlh
    I,t,r.l
    I
    9i9.
    Grcen
    coccoids
    Green
    colonias
    'CTosIeTTopsis
    Charac
    i
    um
    sp.
    A.
    Draun
    sp.
    l
    1
    ipp.'ro*.-Lesn.
    ElggT:am
    !gq!-et!ut
    (Tfiuret
    In Breh) l(om.
    -Lesn.
    lcerreTesmus
    spp.
    lGyen
    Scenedesmus
    bijuqa
    iTurnin)
    LaQerh.
    5cfie-a6srnus
    Tinio-rprrus
    (irrpi
    n) Kutz
    .
    Scenedesmus
    quedFi?audi (rirrpin)
    De
    Bretrisson
    f**
    ryq-rpp:-m}?d
    -_.
    Un
    ili"-nfiTi
    ed
    eug leno
    i
    rJs
    l,
    .....
    l;
    I
    l'.
    I,l
    I
    I
    'a
    t.
    ,
    :
    'l
    I
    I
    I
    Uglqslfa
    granulata (Ehr.)
    Ralfs
    etclotETl
    etclotETl
    t t
    qlomerata
    sEmerata
    BachmannBachmann
    ivct
    uyclotel
    ycloteTE
    oteTTT
    la menegfiTn-I
    meneqhiniana
    rieneofil-nTan a
    Kutz.
    Kut z
    fyc-l
    fycmeTllt
    oteTllt
    D
    p
    s s e
    e
    uAo
    uio
    fiEXTqer
    tfeXToer
    aa
    fvdt
    CyCl otel-Ia
    otel-Ie
    fIeTITqera
    fIeTl-Gera
    TiunTiu'r
    .
    ED-anoZfstus
    5tephanodiscus
    hantzschi
    $:T-
    i
    Grun.
    @loqqos
    fm5er)
    Hasle
    Ol;Toma
    ur a[orna
    vu
    vulgare
    _"_
    tgare Roiy
    uory
    l1-qg_l1
    rraql larra
    qfr a_
    spp.
    snp. LvnohveLyngbye
    tr1ilI1$
    .uj1r1-il-1-u.
    rc:gl.plg
    (Rabh
    .
    )
    Grun.
    e?r-] o n
    eTffiaCTTT urrr
    (
    Gr u n'..)'. rlle r
    e
    s c
    h k
    ow s
    k
    y
    a-----'
    rragl
    i--
    larta
    cr0t0nensts v.
    oreqana 50v.
    rragt
    tdrla
    vaucnerle
    meA-ra
    spp. ETt-
    -
    S:111e0Ta i
    nc
    i
    sa
    Boyer
    @qlq lt:flgl.
    v.
    !rylg.atq
    (Grev.
    )
    Pat.
    Svneilr?
    rump-ens Rr1z.
    ::::::-'
    -
    !/nedrt
    uTna
    lTl
    tzsch)
    rirr.
    fttr-n'anthes
    ex i
    gua
    Grun.
    Achnanthes
    lanceolata De
    Brebisson
    m-nantfieT
    ]ancrTtfa
    v.
    dubia
    Grun.
    Ellq1r!@ flr-rcIg
    (KuT7:I
    cteve.
    Iq1Fn-r3Tpir.
    Bqrv---,
    Navicula sp.
    l9
    mf}_ql._
    q_qp_Uqlg
    v. hungqlica
    (Grun.)
    Ross
    Igy]-.rE
    trvEocephalE
    vl
    veneti
    (Kuti.)
    Grun.
    ilavlc LiiT
    miiiffi.-
    ffitiEuTa
    pT,ilra-cta
    Grun .
    @ne',.@Kutz..
    ':
    :
    E@
    FT---.:-.
    ir_r11uTa
    v.
    .....#
    1e_c_tq4gr4laris
    (Greg.
    )
    Grun.
    Navrcula synmef,rlca Pat.
    Pi-n n u]
    a
    ni
    tn5ls
    c,[iE-f
    r a s s ke
    ]
    'l
    l'
    I
    '1.
    J..t
    '=-,

    -ta
    -
    1.
    --il'
    Scrapirrgs
    from
    an
    entire
    slirJe
    rrerc
    sl,irrerJ in a n-ar-irrg trlentlr]r'ttrrtl
    ,iiiutcrJ
    Lo
    a
    l(n0wn
    volume
    (e.9.,
    '50 ml).
    Oupllcate
    sarilples
    '.Jere
    then
    eounLed
    hiy
    the
    '.arne
    proccdure
    outllned
    for
    phytcplankton
    sarnples
    (Sectl0n
    5,
    Part
    t,
    hcr'cirtl
    pxcript
    that the
    appropriate
    rnultipl
    lcatlon
    f
    trctor's Hcre
    used
    to
    qive
    densit
    ies in
    algal
    tunits
    Fer
    l0
    cmZ.
    RESULTS AI.IO
    rJISCUSSICI{
    COIIPOSITION
    Seventy-five
    ,_*-.J---'
    alqal
    taxa
    were ldentlfied
    from c0l
    iections
    of
    periphyton
    from
    ; expected, nany of
    those
    same'taxa
    were
    found
    in the
    Coffeen
    Lake
    per.iphyton
    col
    lecticns
    {'Section,5,
    Par't
    I,
    herein).
    :
    .
    .:
    ABUNDANCE
    :r
    '',
    Similar
    periphyton
    densities
    were
    present
    at Stations
    .l.5
    and
    2
    rlurinq AuqusL
    1978
    (Tab1e'3.2).
    ,
    The
    periphyton
    Jensities
    decline<j frorn
    roughly
    2,500,:000
    units
    per
    l0
    cn?
    in
    August to
    approxirnately
    .l,200,000
    by November
    l97s
    (Tablt:
    3.2)
    ',,:.i:
    Stations
    I
    and
    4
    lrad
    algal densities
    of apprcximately
    234,000 anct 869,000,
    respectively,
    in February 19i9;
    those
    densities
    were the lowes'trr:ecor,led'for
    the:'
    .
    --.-: .
    _-_
    study
    year
    -*/'--'-
    and-w9_Lg_t1g*J!ejJ_d*L€sul-L_ol_reduce_d__lj.g[_!*a_1g_]libiI
    -.:.::--
    ity
    arrd
    Tan[#-qguie-wtrich-contr_ibulg{lq.ilo1le1-g19ry-t_h,,1a!es.
    _-*;a---
    temperoture,which
    contributed
    to slower
    qrowth
    rat
    The
    densities
    of
    ------l-,---::.^-*-,-v-*--.^"---'"---:
    periphyton
    in May
    I979
    were
    substant'ially
    qreater
    greater
    than those
    recor:ded
    i n
    November
    l978
    and'February
    t'.,
    : ,
    ::.1
    .l979
    at.all stations;
    Station
    I had
    densities
    of
    2t3?6,200,
    and
    Stations
    1.5,'?,
    and'4 had densities of
    3,038,510, 4,144,400,
    ond
    6, 1,48, 360,
    :rrespect'i ve I
    y.
    a
    |
    ,a.
    ,'.
    t,t,
    ,
    ,, ,
    I'he relative
    importance
    of
    the
    major
    stations
    on
    any
    given
    sampl ing
    date,
    dif ferent
    in near.l.y
    every
    col lect ion
    domi nant
    group
    i
    n
    ::the Augnst
    ',1978
    ,'and
    1,.:':.
    '' ..,-.'
    ''
    div'isions
    of
    '
    algae
    was sim-ilar
    between rnost
    but
    the
    most
    abundant alqal division
    was
    (Table
    3.3) .'
    The Baci
    I
    I
    ar
    iophyt,l
    were
    Lhe'l
    May
    1979
    collections,
    the
    Chlorophytd
    were
    =
    =
    :.-{
    j;,ll',:';"".:r
    : :,
    r!.q
    J
    ,1
    3;3
    :,:,::.<

    '-==
    Iable
    3.1.
    Algal
    taxa
    identif
    lerl
    f
    rom
    qirarterly periphytort
    :
    collecti0ns
    frorn
    artificial
    substraLes
    aL
    four
    sampl ing
    ,
    stations-in
    Shoal
    Creek frorn Auqust lgiB
    throrrgh
    ilav
    1979
    (continuert)
    LlfJonei:
    sp. I
    Pmphipleura
    pe'l
    luc
    ida
    Kutz.
    @D_o!ema
    s'pp.
    EaFdh
    Gomphonema
    cf.
    anqustotum
    (Kutz.)
    Rabh.
    mmpnonema
    0l
    I
    vaceum
    (
    L,,nqbve
    )
    Kutz .
    (bmphonema
    parvul
    um
    Kutz.
    FnFfIa
    mifrilfid-TTlse
    & Rabh,
    Fmcr.
    @lgssglq:
    (Kutz.
    )
    Hust.
    firtzscnla
    soD.
    Nitzschia
    spp. l
    flTtEcfiTa
    srr.
    2
    fitffiT
    sD. 4
    fiffiFffi
    'N-iffififfi
    fTTIForgg
    amphibia Grun.
    (t{.
    Smith)
    nTtzsTTTa
    ef.
    -&
    Tqnorata
    Krasske
    I@
    pale_a
    {Kffi)
    tt.
    $nith
    fi-ft-tscTia
    rostel
    I ata Hust.
    ]@lq
    spp.TwpTn
    Itrr-i
    reTi-a anqust
    at a Kut z .
    S[iFeTIT
    oTiTa]euTz.
    RTuegreen
    coffi-i
    d colonies
    Chroococcus
    soo.
    Narlel i
    Chroococcus
    turqidus
    (Kutz.
    ) Naoel i
    l{erJreedia
    qTiuZ
    a-(
    itr r.
    )
    ttaqei
    i
    'ffiil?Eoi:ia
    Merismopedi
    a
    ia;T.=
    tenuissima
    Lemm.
    @
    gg1_"lt.
    r.lenegh.
    L:lg-b-y1
    sp.
    I
    Lvnobva so. 3
    Hust.
    Lylgu_vg
    sp. 3
    Anrb_qqqq
    spp.
    Bory
    3.5

    ::::::::-
    :.::.,,:
    Table
    3.2. Total
    densities
    (alga1
    units
    per
    l0 square cefitimeters)
    '
    -.',
    ,.::.
    -
    of
    :per-iphyton
    collected
    quarterly
    fronr
    artlf
    icial
    ,.
    ,'
    '-.:":'-,
    substrates
    at four
    sampling stations
    in
    Shoal
    Creek
    f rorn
    :,:,
    :.
    :
    ,'
    ,,
    .,
    Augu1t
    1978
    through t'lay
    1979
    (--indicates
    missing
    data).
    St at
    ion
    August
    l978
    November l97B
    Fe'uru ary
    'l
    9
    79 I'tay I
    9
    79
    I
    1.5
    2,
    4
    2r547r000
    2,584,630
    r,rii,zro
    1
    ,246,1
    60
    I,055,1 l0
    233,580
    868, B40
    2,326,200
    3,038,5,|0
    4,
    7 44,
    400
    6,
    I
    ag,359
    :..
    l
    I
    T
    I
    I
    I
    l'
    I
    T
    T
    I
    I
    I
    l,
    f,
    f.

    rSt
    at i
    on
    Gr
    eens
    (
    Ch I
    orophyt
    a
    )
    T_able
    3.3.
    Oenslties
    (al9al
    :,.1
    -
    (in
    p:arenthesis)
    .1.,
    '
    col lect ions
    made
    i978
    through
    Hay
    units
    ;rr:r
    l0
    square
    ccniimeters)
    end
    of
    majr
    r algal
    divisions
    irlent
    if
    ieC
    quartr,rly
    at
    four
    samplintl stations
    191
    .c:
    .
    percent,]ge
    coi4pos
    i
    i
    i
    on
    f
    rom
    per
    i
    frhyton
    in ShoaI frr'ek
    fra:r'r.AugusI
    Di atorns
    Sluegreens
    Cryptomonarls
    (Bactllariophyta)
    (Cyanophyta) (Cryptophyta)
    I
    I
    t
    il
    1.5
    279,42A
    (il.0)
    54
    3,
    790
    {2't
    . o)
    2,21?,760
    (86.
    e
    )
    l,966,o5o
    (/6.r
    )
    3 5,940
    (1.4)
    59, 370
    {2.3)
    0
    {0.0 )
    0
    (0.01
    0
    {
    0.0)
    tl
    t0.0)
    0,
    {
    0.0)
    August
    1
    979
    '0ther
    -*+--*----
    ':
    '
    't.9,280
    rlt n\
    \,.v,
    I 5,420
    /
    If.vir
    n\
    .t!gve[LeL]_9.78
    1.5
    ?
    4
    845,480
    (60.6)
    i
    30,930
    (s8.6)
    537,470
    (5o.e)
    ,
    76,480
    (32.7)
    239,470
    (27.6)
    751,730
    '(32.3)
    4t 5,340
    ,.(13.7)
    422,130
    '(B.e)
    285,280
    (
    4.6)
    308,400
    l\?2,1)
    34i,030
    (27.8)
    3oB,
    3 20
    (.29.2)
    l4l,t00
    iro.l )
    52,650
    ,
    (4.2)
    94,0
    7r")
    ,
    (8.9
    )
    .Bl
    ,/50
    (37.6)
    lB2,l9C
    (21.0)
    40,170
    ,
    (l
    .7)
    54,
    i50
    (t.s)
    ,27
    ,7
    50
    (1.0)
    {:8,7-80
    (1.4)
    0
    '
    (0.0)
    '0
    {0,0)
    :..
    1
    5,4
    ?o
    {1.0)
    30,840
    (I
    .0)
    46,260
    '(
    I
    .0)
    15,420
    (0.0)
    0,2 30
    {l
    ,2)
    5,6 59
    I o 1\
    5,6
    50
    lt,0)
    5,4 20
    (6.6)
    3B,550
    {4.4)
    7/,100
    .
    {3.3)
    46,
    ?
    60
    .('l.5)
    53,
    9
    70
    {l.t
    )
    54, ?00
    t2,s)
    i0
    Ie!rqqly--1-ei9.
    53,930
    (23.1
    )
    :
    408,630
    (4
    z. o;
    Igv_l_9zg
    1
    ,441 ,
    /80
    (62.0)
    2,490,320
    :(82.0)
    4, I
    g4,2go
    .
    (88.4)
    :
    5,605,1
    80
    .
    :
    ,
    (91
    .2)
    :;:j;l:

    -j:::'li
    i.i:,=
    :r:::
    :-ii:l',;-:r.,
    l.
    lr::=r,:.::l::;
    5
    :"rt..
    ,,-.'
    r:-.
    '
    :i:
    =i:.
    r
    '::.
    I
    llr-:
    l
    .:
    ,'.
    ,;:':'
    t
    :
    t.r..
    l'
    :::
    l.
    t
    I
    t
    t
    I
    I,
    I.',
    :'.
    ,i.
    r..-
    I,:,
    .
    :i:r
    .
    l'.'
    -'.
    ..?.
    l...
    I,:
    l'
    i',
    r,,:
    r''t
    l:l
    :1.'.:
    -::!:,
    I]
    r..'
    .:::ii:
    l':
    do'ninant
    in''Novemtier
    1978,
    and tlre
    Cyanophyla
    nnrJ
    Bacj'l1.tr'irtphyta-wer-e
    Lirtl
    rlominant
    algal
    'groups'in
    the February
    1979
    col lection.
    ;tri,-r,,.das
    spil
    l,*.i.,
    overf
    low
    ftrorn
    the'lake
    curing
    Apri'l
    anri
    :ttay
    tgis but no rnajor-
    ,Jif
    ferences
    iI
    algal
    divisions
    were
    noted
    on
    Euhstrates
    located
    at
    t.he
    sLaLiorrs rrpsLrerlrn
    ancl
    downstr:eam
    frotn
    the
    junclion
    of Lhe
    purnphoilse
    candi
    ,*i'tir
    Shc;t'l
    [resi('at
    that
    tirne.
    DIVERSITy
    '.1
    .
    dnd,'are'presented
    in
    Table
    3..4.'rThe rtiversity-indices
    were
    highest
    i'n August
    lgiB
    (i.e,,
    4.46 at
    Station
    ,|.5
    anrj 4.2.|
    at Stalion
    ?)
    and
    sornewlrdt lower
    irr
    ilovernher
    l97B
    (i.e.,
    2.73
    al Station
    1.5
    ,
    3,03 a[ Station
    Z
    and
    3.26 at Station
    4)
    in Shoal
    Creek, The
    diversities
    calculater!
    for
    the
    f.ebruary
    lg79 collect,ion
    were
    similar
    to
    and slightly
    higher
    than
    Lhosc
    in
    Novernber fg/8
    (i.e.,
    3.03 al
    tl.i.,
    ,,1
    .Statlon,
    1
    a1d 3.a5
    at Station
    4) but
    stil l
    lower
    than lhe
    values
    cal'cirld.-t€ri,'f
    or
    the
    August
    l97B
    collection.
    in
    t'lay
    1979,
    Station
    4
    had
    a
    diversity
    inrlex
    of
    4.41
    and Stations
    1, 1.5,
    ancl
    Z harl vaiues
    of
    3.93,
    3.61
    ,
    anrl
    3,15,
    respectively.
    Throughout
    the
    year
    Station 4 lraC the
    highest
    diversities
    dur-ing
    the
    year;
    the
    reasons for
    this
    trend
    were rrot
    clear hut
    the locat,ir:n
    of Stations
    ,
    .|.5
    ancl
    2
    in
    a
    pool
    formed
    near
    the
    junction
    of the
    pumphouse
    cdn,ll and Shoal
    '
    C:"eek
    may
    have
    influenced
    colonizat.ion
    at,
    those
    stations.
    During
    Lhe
    April to
    '
    May
    exposure
    period,
    the
    lowest
    inttex
    was.6'lqrrlaterJ
    for
    Station
    2.
    'ihis
    period
    was
    of
    particu'lar
    interest
    because overflow
    frorn
    the
    spil
    lway
    occurred
    then
    and
    ..,'',''
    ry?l:!
    mqv
    trqvg:!99-0-leduceil
    bv.-Lhe--q.gli]p-u-gt-l-e!tgG;l
    it
    was not
    low
    enuugh
    "
    to
    i,
    indic_a!e_l_q_t::e5_q*s_i!gatfp.n._and
    .;.--.;.:,-_.-.1._...--__:*:_-._lJ
    the
    much
    hi
    gher
    ;-.+.:4*''.*.il-
    diversity indei
    calculated
    for Station
    4 rJuring
    the
    sanre
    perioo
    inoicalic
    ttlot
    o
    --*
    -.--:-...>_*
    -.-{.,*-*!-*
    very
    diverse
    periphyton
    community
    existed
    only,'a
    short
    distance
    rjorrnstream.
    -
    ,
    r:.'

    F;;-
    i'
    fi
    1.,'
    ii,',
    Table
    3.4. 0iversity
    !ndices
    calculated
    for'per,iplr-vton
    collected
    afl"er
    ,:..
    ,,
    quarterly
    ?tl-day
    exposure on
    gl;lr
    -sLidq-{
    ;rt
    f
    r)ur 51if,ioc!
    .,
    :
    in
    Shoal,
    Creek f rorn
    August
    l9l3
    lhrrrvglr
    4a7 i979
    {----
    Indicates
    missing
    rlata).
    Stat
    ions
    August
    I
    978
    November
    'l
    9
    78
    Februar:y l9
    ?9
    M,ry
    I
    9
    79
    I
    .,
    1.5
    2,7,7
    :],45
    3:93
    3.61
    l.
    I5
    4.
    41,
    ?
    4
    4.
    46
    4,?l
    .:
    ?,7
    j
    3.03
    3.26
    il,
    i
    .:,,
    T
    !;.
    il'
    t:,.
    .','
    ,1,
    iT
    frl
    r$
    .:..
    ':
    3.9

    3
    ..t
    FI,
    =it
    ,
    S[i.I
    irili
    .i
    ]
    Bti{fHIC
    lllYtSIiCAilCX5
    AI
    3,i*;'.i- ;1,,1'
    iry
    .:
    ,,
    ',
    .
    :::
    A
    study,'of
    thc
    benthos
    cf Shoal
    Creek
    vtas
    conrJucted
    ir
    crd€r-
    tr::.:s:rsi rrir
    ef fect
    of
    heated
    water
    overf
    low
    f
    rom
    Coffeerr
    Lake
    rpcri
    the b,rnlri'ic
    f r,.rrie.
    Ihr-ft
    stations (one
    control,
    two
    downstrea,n
    f ro,n inpacr.)
    *,1.."
    sal;rli-c
    .ri,rr'
    I
    grre-ygr.
    periOd.
    The
    taxOnr,.,liC
    CompgSitionS
    of
    tnc
    thrl'p
    lOCrtions
    ;re1t
    :i;:ri
    lJr-,
    lrut
    differences
    ilere
    present
    in rJensities
    of
    tlre
    do,nins6ll
    slQ3nis,,r5
    6rn,li1
    r.
    llil
    ttlrrJ,j
    stations.
    Al
    I
    co'llections
    were
    rlorninatetj
    by
    ilrn,l[,rre:ili
    ilre
    h,:ctl
    anii
    t,re
    r:ii,j!ri
    gerlera
    Chirorlgmus
    and Polypedi
    I un. Analy|*s_
    ol__i_g:'ip.gtl!v3_r..nod_jjssirl:g,1-_.:r.Iy;jt_:n
    data recorded
    congu.rlent.ly,ni.Ln_benthjc_co_ll-SJ!i.qnS_.irUJi-r,,:!_eJ
    .!jr-A1..-jgrq!,1_.-!{
    thesr:
    two
    oarameteLt-l{g!:e.
    r{j-!"lUn
    -.-;.;:-:_.
    -t_hg.tgle.r__ance
    -'tiniLS
    ,lf
    _:;i.:sL
    lrenttrir-
    LJr.J
    dur,irtg
    !:-.
    _!_*_.-t:t9--9ygf
    f.Lq!-
    p-er:-i-o-ds,.a.l.
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    Lhe
    ucnthic
    cclir:i,i.i*ns.'l
    aL
    al I stotions
    anrl
    accounterl
    for
    501 of
    thr: total
    orqrrnis,ns collact.r-,tl.
    Itrc
    0llgocltoeta and,Chironornus
    pref
    er 0rgnnlcal1-y
    enriclrcri
    habii.atr
    Hherr'ille
    "
    :
    ol
    igochaetes rlerive
    their nrrtrition
    f rom
    bacteria
    and
    Cnironcnlur
    nre |
    /ie-
    truilding
    detritivores.
    Pql-yped_ilunr havc
    been
    founrJ in
    a
    variety uf
    habitaLs
    anrl
    have
    feeding habits
    ranging frorn
    preclatory
    to
    deLritivorous.
    r)ther
    irnportanl"
    taxa
    occurrinq
    in
    Shoal
    Crr:ek were ilre midges
    &!1|q1s,
    Dicrotq4igq!, on,t
    ,proqla{iu!,
    anrt
    the nrayfly
    caenis,
    iU.rtj:lelj_qlq*Ugig1]-rglt::_:_?1g"
    'oiruun
    i !'ies
    '
    .:.
    DIVERSITY
    ./
    !
    Oiversity
    indices attempt to
    surnmarize
    corrynuniLy structrrre
    s0
    LhaL Ihe habiLat
    \
    (.
    /
    and
    water
    qual
    ities
    of
    'tJif
    t'erent
    Sampl
    ing
    locations
    rnay
    br:
    cornpared. ln the
    diversity
    index
    proposerl
    by
    Shannon
    (1948),
    values
    of
    less than I
    irrdjcaLe
    he,rvy
    po1.lution,valuesoflto3inrJicatemoderatepo|lrrLion,anrlV,.}lt,eSabove3:..
    denote
    clean
    iwater
    areas. Div,ersity indices
    are nosl
    acc,traLe:when
    al
    I t'axa
    :
    .iested
    are
    identified to
    species;
    llowever,
    due to the
    poor
    taxonornic
    state.of
    Inany
    aqrlatic
    qroups,
    iderttifications
    to Lhis
    lr:vel are often
    inrpossillle.
    in
    sucit
    situatiorts tJiversity
    inrlices
    lose
    sorne
    of
    tlreir accuracy
    and
    rnearring.
    0iversiLyva1uesoverthecourseof.:theShoalCreekstudyrangedfroinl.37.
    1St-ation
    4,
    January,
    1979)
    to
    lf
    .aS
    15tation.'l,"May
    1979).
    Seasonal f luc.+.'uaLions
    l
    were very:sjrnilar
    at the three sarnpling
    locations
    (Fig.
    4,4),
    I'lo
    one
    sta-l.ion_-,
    w4ljle4rJy nxue-.q1-.l9f9--q-lVe:9.-e,-.t!-O1l-q.l.y
    o-t.b-qf ovel
    !!e
    qo-,l1,re
    ,rf-
    [i-
    iilJv
    .
    pq',:gd,
    A value'0f 3.85
    rvas
    calculaterl fo'r the
    control
    stat;lon nci'' ihu.;ir
    or
    lfte
    faxe overflou
    period
    i'n
    l4ay
    1979,
    while
    lower values
    were
    calcul,rted
    for
    :
    :
    Stations
    2
    and
    4
    during
    this
    same time
    pr:riod
    (3.64
    anrl
    3.j8,
    respectively).
    However,
    the fact
    that the
    values calculated
    for
    all stations
    were
    \{elI
    abovr:
    3.0
    dqr11g_this
    tjr!s-ira,ne-jndi@.!,i!{
    there
    was
    I
    ittle
    itr$;ixr-ix:[t1iiC-::*
    ]l
    :,..;;*-*-':-''
    habitats.-
    _-,
    :
    ..,
    i.
    ,
    :
    .,
    i
    1,,
    l'
    ,tr.'
    .t
    ::"::
    :
    .
    :
    .:.,
    :
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    ::
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    .lj
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    ...,;-
    't','
    'l
    ili.
    :.-#
    I
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    :.:
    ir.
    ta::::
    ...

    il
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    ;r.l
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    ifarft
    0ltlllt'l
    ad,
    .ut
    tlrf
    !!q,!l
    c4ltxfl.nr!
    ".r:,
    iil:!,r
    ltrl.'(:t
    tlrl!lor':t,glrda.i
    (Atti,
    rotlhltlri
    ltfl
    ii4tri!::-:r
    :t'
    i!!!"i
    !
    |
    '
    i.
    ft
    R
    f,
    n
    lfiqlt
    I tco
    I
    rr$0
    I r40
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    teo
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    rco
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    CFflx.
    itltHBt^
    t'179
    t'e4,!r/t
    'it'
    rt,".r-
    r.;.r
    l'i:r
    :
    '
    {lnil
    trt|stit
    nl
    li!(
    |
    Ivl
    rt"l
    tlrrdlotrit
    lttrn't];lL
    ltn
    rr'rlrt
    5lnl
    tOrl
    {, sllonl
    fRftX;
    il4vr{8t n
    l9r3
    lrrer!'lfir
    .lr!l'
    'rtr
    il
    t t
    I
    r
    n
    d
    fi
    0
    5
    I
    OO0
    EO0
    600
    a00
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    260n
    I 400
    2?Ott
    20n0
    I
    Oflo
    I
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    I ao0
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    f,00
    :
    atr{)
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    0
    ''::
    .'
    ,:-
    .
    i
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    .
    '
    :
    [+Vtf'
    MCan
    densi
    ty.of
    ot-'Shoal
    .
    Creek
    ,ceptembeF
    1979
    .
    rrl!?1
    tsn(
    ,1
    rn'
    ;1
    the
    .f
    i
    ve
    :nlos
    t
    Stations
    l,
    2,,
    n
    f
    n
    ta
    tl
    n
    l{
    0
    I
    .l
    abundant taxa
    .collected
    fronr
    each
    and
    4
    from
    Novenrber
    l97B
    through
    ,ii::+'
    ::.::.':.:a:.:
    4rl

    ':.
    1::-:::
    I
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    l.'
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    I];
    r
    'tl
    T.
    $
    l=
    lJ
    :.4;
    '
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    ::,'.,::
    :::.
    ,:.:,
    i.:
    :,i
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    a)
    rlJ
    t/)
    '0J
    qJ
    .L
    :,{o
    \ft
    t*
    <>
    V,
    .1,
    1. o,.
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    ar<
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    ra-
    oJ
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    q.)
    cJ
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    4.8

    :.i,
    I
    LlltRAT|ll{t
    lT
    ctI[0
    t
    .
    ':
    ,l
    t.,
    Curry,
    1..-
    [a
    1965. A
    survey
    of
    environmental
    requirernents
    for Lhe
    rnidgcs.
    ,ln
    9., T:
    Ta.rzwel
    l,
    ed.
    Qio!ogical
    Probletns
    in"diter
    Polluf
    ion, Thircl-srminiF,
    Publ
    lc
    tlealth
    Serv.
    publ
    .
    999-il0-ZS.
    pp.
    127-l a.$.
    '
    ,'
    .
    Gauf
    ,'
    in,
    insects
    A,.
    R.
    to
    and
    heated
    ,s:
    Frn:
    waters,
    l9il.
    Jour.
    raboratory
    .Kansas
    tntornol,
    studies
    on
    soc.
    toler.ance
    4itzqo--2is,'
    of',
    aquaLic
    '
    :-
    t'
    Hubbard,.M:.D.
    and
    lJ.
    L.
    Peters.
    l97B. [nvironmental
    requir.ements
    anrt
    q9l
    lytion.tolera!!g
    gf_Ephemeroptera.
    Environ.
    Hon,
    'supp.
    Lob.
    ,
    ustpA,.
    Cincinnati,
    0H
    (iPA-600/4-7S-061
    ).
    46t
    pp
    Larimore,
    R.
    .|970,
    Two
    shallow-water
    bottom
    samplers.
    prog.
    Fislr,
    cult.
    l
    32(2):l
    l6-l
    19.
    l{ebeker,
    A.
    V. and
    A. E.
    Lernke.
    .
    .l968.
    prel
    iminary
    studics
    0n
    the
    tolerance
    of
    l
    aquatic
    insects
    to
    heated
    waters; Jour,
    Xansas
    Entonpl.
    5oc.
    4l:413-4.l8.
    1:
    Shannon,.
    C.
    -E.
    1948.
    A
    mathematical
    theory
    of
    corrmunication.
    Bell
    System
    Tech.
    Jour.
    ?-7:479-423,
    6?3-696.
    ',.:l
    -
    .:!:.::.;,
    .rl::
    i=
    .
    4.9

    ,
    ,
    -,'
    j
    :
    :
    StcIlDt{
    -"-'-'-
    5
    '
    'a
    ,
    TFFECI.OT'
    A
    IHER},IAL
    OISCI{ARGI
    Oil
    r
    I SHI5
    OT SHOAL CREEK
    ,
    bv
    ,'i
    rtt
    Lance
    G, Perry
    .
    ABSTRACT
    ntr., I t\t1u
    I
    An
    t-
    -.
    investigation
    -
    of fish
    distribution,
    abundance,
    and species
    composition
    in the
    East
    Fork
    of Shoal
    Creek
    was initiated
    in
    September
    19iB in an
    effort
    to
    assess
    the
    impact
    of thermal
    enrichment
    on
    the fish
    community.
    Species
    composition
    of
    the
    East
    Fork
    ichthyof
    auna
    was found
    to
    be
    sini'lar to.that r-eported
    previously
    f
    rom
    the
    area.
    -
    Althotlgh
    :lltltc_jgt
    L-e-q,pg"fglyry_e_lg-y_qtigns
    lej-e*1-a1e-
    dur
    jng
    the
    __slu_d_I.r-!!9*plg!9!lce!
    lherma!_infIugtfql.9.r*_:pg:jel__qi*Ligyt_rql_g1_glgndance
    .
    '...'
    .
    were
    detected.
    '
    Anticipatetl
    occurrences
    of
    tneimoi,
    ;;;i;h;un-t
    in
    the
    creek
    (pre-
    .
    ::.':
    r,mably
    durinq
    the spring)
    probably
    wou'ld
    not
    exert,a
    lethal impact
    upon exposetl
    fishes,
    but
    may alter time
    and/or
    duration
    of
    spawning
    activities
    and
    rate
    of
    development
    rluring
    early
    life.
    It is unlikely,
    however,
    that
    the
    sur,rival
    capa-
    city
    of the
    strearn
    populations
    would be
    jeoparrJized
    unless
    critical spawning
    grounds
    were
    regularly
    'inundated
    with
    excess
    heat.
    A
    worst-c,rs,:
    l,nidsummer)
    occurrence
    of thermal
    enrichment
    may elevate water
    ternperatures
    to
    lethal
    ,:
    Ievels,
    but
    such
    an event
    presumably
    would represent
    a
    temporary
    and
    localizerl
    '
    area.
    a
    ="-
    '-:-
    ;.-t-/
    -IT
    :
    -'
    .'
    -!
    '.
    :-..:a.
    5.1

    :::
    li
    I.:
    .t
    l',
    !
    ':'
    :.,
    :
    :
    :
    ,
    .'l
    Coffeen
    Lake
    was
    constructed
    to
    serve as a
    cooling w.rLer f
    acili
    L.y
    f
    rsr
    Cerrl.rnl
    Il
    l'inois'
    Publ
    ic
    fur'vlce
    Company's
    Cof
    f
    cen Power
    Station.
    Sincc
    cl0sur"e
    of
    the'
    dam
    w-as completed,
    impounding f4cOavid
    Branch,
    iln intermittent
    Lributary
    of
    Shoal
    Creek's
    East
    Fork, the
    potenLial
    has
    existed
    for
    occasional
    lherrnal
    enrichment
    of
    the
    area
    imnediately
    below
    the
    dam.
    Frequency
    of occrrrrence, magnitucie,
    and
    rluration
    of thermal
    loading
    in
    the
    East Fork
    cannot be
    predicted
    with
    certainty
    since'those
    events
    that
    cortlrl
    produce
    an increase
    in'water
    temperature
    0ccur
    at
    irrcgular
    intervals.
    Tlre
    release
    of
    thermal
    effluents
    from
    the
    lake
    is
    ultimately
    regulated
    by
    water
    level;
    it
    must
    erceed'590
    ft in
    elevation
    (above
    me'an
    sea:.level
    )
    in
    order
    to
    establ
    jsh
    and
    flaintain
    a
    cJisclrarge
    over the
    spillway.
    Beyond
    that
    initial
    requirement,
    the
    cdpacity
    of
    heaLcd
    lake
    water to
    induce
    ternperature
    elevations
    in'the
    creek is
    contingent
    upon
    power
    pl:ant
    operation(arnuntofelectricitybeinggenera|er!),andambientwater
    ternperatures,factorsthattypical.lyvary.onaseasona.lbas.is.
    Althouqh
    the biological
    impacts
    of
    periodic
    adrjitions
    of
    excess heat
    in
    Shoal
    Creek
    are nutnerous
    and
    potentially
    significant,
    no
    cornprehensive
    jnvestigations
    aimerl
    at:evaluating
    thermal
    effects
    have been
    corrducted
    in the
    l6
    year-s
    since
    l
    impoundment.Anopportunitywasprov.ided,however,inassociationwiththe
    Coffeen
    Lake studies,
    to determine
    species
    composition,
    djstribution,
    and rela-
    tive
    abundance
    of
    fishes
    inhabiting
    reaches
    that
    are occasionally
    subjected
    to
    therrnalenrichrnent.Theprilnaryobjectivesofth.isstuc|ywere.to,]c0nrparethe
    above-stated
    parameters
    at amllient
    and thermally
    influenced
    locations
    and
    to
    relate
    those findings
    to
    published
    accounts
    of
    past
    fisheries
    surveys in
    order
    to
    note any
    changes
    in
    the
    local ichthyofauna
    that
    may
    have
    occurred
    since
    impoundrnent.
    METHODS
    AND MATIRIALS
    ,
    t
    :
    Stations
    1,2,3, and
    4
    (Fiqure
    1.1,
    Section
    l,
    herein)
    "t'
    November
    1978, and
    on
    5 Apri 1,,
    lB
    May,
    anrl
    20
    .luly
    1979.
    i
    n an
    area unaf fected
    by
    therrna'l
    d ischarges
    f rorn
    the
    I ake
    ,
    -'andl4
    ieceiverl
    occasional
    ther.mal
    inputs'.
    Sarnpl
    ing
    at
    ,-
    t
    I
    l.
    a
    t
    I
    I
    ':,
    I
    I
    l'
    ,l'
    I
    l
    I
    Fishes
    were sarnpled at
    on ?B September and
    30
    '
    Station
    I
    was located
    r,ihereas
    Stations
    2,
    3,
    ::.':',
    ,:
    {::=',;t,..;+,-

    ::*E
    .a:{
    =
    :'r
    itt,'
    ]'
    i::r.i
    .r
    l,t
    Iir
    I,.'
    ;j'-1
    ,
    1,.:.
    ,,;1:-;;:
    ,
    '-
    i:::
    .
    .I:
    .I
    il
    I
    edch
    Stdtion cOnsisted
    of
    one
    barJ srtirrri lraul
    of l5
    i.,'rtr:r
    Cigt",rncr:
    in
    d
    'rJoitrt-
    stt:s6ft1 directi'on.
    lt
    was
    origin0l ly inten<tr:'J
    to rtmpil)
    eacli
    slaLir:rr
    gvrir.y
    Dt.h'lr
    mOnth
    (tlirnonthly)
    but
    dn
    ice
    cover
    in ,)dnuary
    anri
    tt*rcir
    [)rr]clu{,'lrl
    cr;llrr.t,
    in.1
    at.-
    t.ernpt's
    dur
    i
    ng
    ,tlrose
    months
    .
    '
    '
    ,
    A1
    |
    f
    ishes
    were
    preserved
    irr i0S formal
    in
    ,:nd
    irJentiiie<J Lo
    5pecies
    {eriept
    hybrid
    sunfishes) in the laboratory,
    ldentifications
    were
    dcconrpl
    ished
    witlr
    tne
    aid
    of Smith
    (I979)
    and that
    refererrce
    also served
    as
    a sor,r-ce of
    information
    on
    the
    prior
    status
    of
    the
    Shoal
    Creek
    fish
    ccnrnunity. kientific and colnflon
    na$es
    of
    fishes
    used herein follow
    Bailey
    et. al.
    (1970).
    RESULTS
    ANO
    DISCUSSIOI'{
    f ish
    col lecting
    efforts
    in
    Shoal'
    Creek
    yielrled
    a total
    of
    915
    irrdividuals
    repre'-
    senting
    j-9"Y9Jt--f-anil-tef
    --qtd.4
    spp*qlps
    (excludirrg
    hybr-id
    sunf
    ishes),
    Minnows
    .,
    (Fami
    ly
    Cyprinidae)
    were representecJ
    by
    9
    spr:cies arrcl
    were of
    greatest
    irnpor-
    tance
    numerically,
    comprising.
    7ll"
    of
    the total
    catch. 0ther taxa, in descencling
    order
    of numerjcal
    importance, inciuded
    sunfishes
    (Cent.rarchirlae,
    5 specics antl
    l5% of total
    catch),
    topminnows
    (Cyprinodontirlae,
    I
    species
    anrl B%), suckers,
    (Catostomidae,
    3
    species
    and
    2%),
    darters
    (Perci,lac,
    2
    sllecies
    and 21),
    shad
    (Clupe'idae,
    I species
    and
    21(),
    and
    catf ishes
    (lctaluritlae,
    I species
    anrJ
    0.21\.
    Species
    encountered rnost
    frequently
    includecl Llra red
    shiner
    (Notropis
    lutrensis,
    32% of
    total
    cat'ch), sancl shiner
    ({.
    ltlamineus,
    i4f), bluntn*iili'o;
    .,,,
    rochirus,
    l0%).
    The assemblage,iof fishes found
    in
    the
    study area
    appeared to be
    typical'of
    tlre
    reqion based
    upon
    earlier
    reports.
    Df 23 species
    known frorn
    Lhe area
    (Smith
    1979), 13 were
    encountered
    in
    the
    present,
    study.
    Four
    species that were
    repre-
    sented
    in our
    col lections, including the
    0olden
    shiner
    (tloternigonus
    cryso-
    leucgs),
    bigmouth
    buf
    f
    alo
    (!*j-g!uJ
    cyprinnelus),
    qqi
    I tback
    (Carpiodes
    cyrpinus),
    and black
    bullhead
    (lctalurus
    melas),
    were
    not
    found
    previously
    in
    the immediate
    study
    area
    but
    were recorded
    frorn
    adjacent
    localities
    witJrin tlre
    ,
    decirnated, rare,
    or restrictetl
    jrt
    nor was their
    presence
    clocumenLed
    Eait.,Fork
    systern.
    Fishes
    designated
    as
    either
    Illinois by Smith
    (1979)
    were
    not encountered
    by'pr.evious
    invest
    igatois.
    :l
    I
    1I::
    ti:-
    ::
    ,1.'
    .I
    .Il
    1..t,
    :::l
    il
    i=:t,'.t.
    1'Pr:l
    '.
    !
    l:.1
    i:r I
    !i:1:i
    :aii
    i::::
    ..
    -.:
    a:4.
    ,i!:
    5.3

    No outstanding dif ferences in
    speciRs'Cornp*:ilion, i€inpr:;ral ociurr"elrc0, ot' abun-
    ff
    dance of.
    individuals
    were
    dete
    in a
    cornparisan
    rrf,Lhc
    ar*ri*nl. sL,ltir.in
    '*ilh
    therrnal
    ly-inf
    _
    luencert
    :- _--.-
    staLions.
    Stauoni-f
    llif{Entl-;
    il;;;-
    j-r,icrr
    }tc1"i:rr
    i
    total
    catch
    cornprised
    of six families
    and
    l4-ll
    species,
    antJ tnr-'St,rtiorr
    4
    caLcn
    inc:lurled
    leven
    f ami
    lies an<J
    17
    species. Those f
    inrlin^rts
    tlo
    not. nccessarily
    reflect
    therrnal influences
    in thc
    craek,
    llor{ever.,
    sirrce
    the inpuL
    of
    heaterl
    lake
    water
    was not continuous
    throurtliout
    the
    period
    cf sti.tdy. Snow
    nrelt arrrt spring
    rainsma.intainerilakeleve]ssufficientlyhightosustainanirttermittent
    discharge over the
    spillway
    from
    early
    l4arch to nrid-l4ay
    (i979)
    only.
    Theref
    ore,
    of f
    jve'monthly
    sarnples
    obtained
    during the
    past
    yearr
    onl.y
    tlre rnonth
    of
    'April
    representert a
    period
    of
    potential
    thernral impact
    in Lhe stucly area.
    Tht-.
    ternperature
    of the discharged
    effluent
    on
    the ApriI
    sd'npling
    date
    was ?7'C
    at.
    the time,of
    release
    from the
    plant
    (J..1-.Kerrnedy,
    person,ll
    comniunicaLicn).
    Recorded
    water temperatures,at Shoal
    Creek
    sarnpling
    stations
    were rnuch
    lower
    ,
    however: Statjon
    I
    (ambient)
    =
    8.2'C,
    StaLion 2
    =
    10.9'C,
    Station
    3
    =
    9.5oC,
    and Station 4
    =
    9.5'C.
    ll!I$j:t.r,lq'ipgr.-utures*aL-lesL*slalionS:wrl-*:
    -._:
    -,x
    sliqhtly
    sufficient
    F
    hiqher
    magnitude
    than ambient.
    to
    producq_a
    it
    is
    noliceable
    douhtful
    /--r-
    djsparity
    that the
    observed
    in distr'ibutions
    irrcrcases
    ofwerc
    of
    -
    -:--.-*:'-:---."'".-..-_.
    fishes
    lletween
    heated and ambient sites.
    That
    was supported by our.findings in
    +,--
    l'-
    iFeffifreTtri I co'llections
    were simi lar
    in
    species cornposit
    jon
    and
    aburrdarrc.r) <it,
    the four stations. In vier,r of
    the
    prel
    irni rtary
    anrl somevrhat
    ctrrsory nature of
    .
    those
    collecfions,
    however,
    the data
    were considered
    inarlequate
    for
    rnakinq
    any
    '
    def initive
    judgernents
    on
    tlre generai
    wel
    I
    -being
    of
    Shoal
    Creek
    f ishes
    or':
    ,
    ':i'--
    ,"
    uti.l
    ,
    i
    ty
    ;to
    review
    l
    the basic
    -'ecological.
    requirenrents
    of
    the
    local
    iclrtlryof
    auna
    'i
    rr
    l'.order"tb
    '
    :.
    allow a
    predjction,
    bisecl upon inferential
    evidence,
    of
    chaqges,'in the
    ,,
    fish community
    that
    may
    have been,
    produced
    by
    thermal enrichrnent.
    The
    reqrainder
    ,
    ,'
    ::'
    :
    '
    '
    ..of.thisi..reportisdevoterltothat€itd...
    Any rnodification
    of the
    natural
    thermal
    regime
    in aqrratic
    systerns may
    signif
    i-
    l:-.
    cantly
    alter normal
    physiologica'l
    and
    behavioral
    furictioning
    of exposert fishes.
    ..,,.'
    Because,f ishes
    in
    temperat,e regions
    have adapted:
    to a rather strinqent
    dr1il-,'.3l
    .
    :,.:.,-..
    ,
    ,.'
    -,:
    ,,
    .
    l
    .
    thermal
    ,regime,
    and since
    their
    metabo'lic
    rates .are djrectly
    related.lo:wirter
    .l
    temperature,
    daily
    activities and
    biorhythrnic cycles
    (rnigrations,
    repr'oducLive
    timi.ng, etc.)
    are,often
    keye.d.
    to :seasonal temper'ature cycles.
    ,'Accordirrgl
    v,
    :.,.,:.:dFt,i:f
    .'.'
    icial.tempeiitur"
    '.
    areuationsrmay-"'affect
    f
    islres
    'in
    lrn,erous,
    diversl,''
    und
    T
    I
    I
    I
    I
    t
    I
    I
    I
    I
    T
    t
    I
    a
    I

    ,l
    i
    a,
    ,l
    il
    :1.
    il
    'f
    :.-
    t
    :l
    il
    il
    il
    a,':i
    ;iii:l
    ]iiJ
    fl
    '.
    'l'
    complex
    ways.
    l'he
    comptexity is evittencer! by a
    mulLipligily.of
    possiblr
    inter-l
    actions
    involving
    lethal and
    sublethal
    temper,rture
    ef
    f
    t:cts on Lilc
    one,h,lnr-1,
    and,
    varying
    rtegrees
    of
    temperature
    sensitivities
    exhilrited
    try the
    various
    lii,,
    stages
    (egg,
    larval,
    juveni)e,
    arrrl
    adult)
    on
    Lhe
    other.
    A
    irnowlerlgc
    of
    life
    history
    strategics,
    habitat requirements, anrl
    tenperalure
    tolerances
    of
    Shoal
    Creek,
    fishes
    is
    required,
    therefore,
    lrefore
    infere,ices
    cdn be
    rJrayrn regarding'
    their'vulnerabil
    ity to
    therrnal impacts.
    A
    sumrnation of those
    daLa, derived
    from
    the
    literature, is
    presented
    in
    Tables
    5.1 anrl 5.2, and the following discussion
    is
    irr
    reference
    to those finclings
    ln
    general,
    fishes
    exhibiting the lowest tenrperature
    tolerances
    also
    exhibit
    the
    earliest and
    possibly
    the
    shortest spawning seasons,
    In
    Shoal Creek, this in-
    cludes
    the
    suckers,
    darters,
    and
    among
    the minnows,
    the
    creek
    chub
    (SgrnoLilu!
    atromaculatus).
    Four of
    those
    species, the
    creek chub,
    creek chubsucker
    (Erimyzon
    oblOnqus_),
    white s,lcker
    (Cgt_o:!.qinys
    cornnrgrj;oni
    ),
    anrl
    johnny
    darLer
    (E!!eo{omu.
    niqrum) are characteristic
    of
    Pfleiger's
    (197
    l) 0zark and
    0zark
    prairie
    faunaJ regions ancl, as such,
    probably
    require
    a
    clear watcr
    arrd rocky
    bottom
    habitat for long-term
    existence.
    Clear
    water
    appedrs
    to he
    thc
    principal
    reqoirernent
    of
    the
    slough darter:
    (Etheostgnra,
    gaclg)
    as wel l,
    an
    irrhalri
    tat of
    Pf leiger's
    (1971)
    'lowland
    f
    aunal
    region. The remaining
    nembers
    of those f arnilio
    groups
    typify
    either
    tlte
    prairie
    assemblage
    (qui
    llback
    arrd
    blacksirJe
    <larLer,
    Percina
    macul
    a!a), usual
    ly
    inhabiting
    nroderate
    ly low
    gr.rclient,
    w,trrrwater
    ;;*t,
    ;r
    td
    "wide
    ranging"
    group
    (bigrnorrtlr
    buf f
    alo)
    whiclr
    are bel
    ieved
    Lo
    exhibit
    a relatively broader
    range of
    tolerances for
    water tenrperature
    extrentes,
    turbidity,
    level of
    dissolved
    oxygen, volunre of flow, and
    subsLrate
    cornposition
    than the
    aforemerrtioned
    species.
    As
    a
    group,
    the rninnows
    dppear
    to
    possess
    a
    greater
    tolerance for high
    water
    tenrperatures than
    Lhe
    suckers
    or clarters
    (Tatr'le
    5,2:l';,'l'lost
    are
    spring
    and
    early'surrmer spawners and
    a
    fevr
    have
    spawninJ
    seds0ns
    extending
    into
    late
    summer. A wirle range
    of
    envjrorrnrcntal
    tolerances
    is
    suggested
    by their designation
    as
    either
    prairie
    strcarn inhabitants
    or
    ::
    :
    wide-ranging
    ,in
    distribution
    (Table
    5.1).,. Two species,
    tlre si lverjaw
    nrjrrnow
    (gfig!.g
    buccata)
    and stoneroller
    (Carnpostoma
    anorntllurn)
    apparently
    have more
    stringent habitat reguirements than the
    others
    arrrl
    tlisplay
    a
    greaLer
    aFfinity
    for
    clear
    '
    water and
    rocky
    substrates. The gizzarrl
    shad
    (Dorosnrrra
    .ggffgj-U!,n),
    bl'dbk.'bullhead, blackstripe
    topminnow
    (Fundulrrs
    rrotatus),
    and
    the
    suhfishes
    a'll
    5.5
    :

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    {eoerally
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    '
    re,ndin.riLhin
    thg
    confines
    of the
    ldke.
    llonr:th1'1i,.,r,
    .1
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    .,' of
    tlre trff:ects
    of
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    :lrel:
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    strorrl;l
    !rlr'lrldr
    oll
    pi;?r:n!
    r
    rl
    Inpacts
    assoclatcd
    with the
    occurrencLi
    rif
    a worst
    -r..r:rr
    tir,:r,ir!
    1,{orJ.,.j/r.. .i!i
    I
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    :..orre.1al:d!.an'Jdmoret.ypical,pr-artictabiel,ve|tLrrnl.itr:4il1*f
    ::,:
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    -rrorst
    -case
    Ec(
    dlscharge
    becduse,,
    .A:
    ndr
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    !y
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    .J1.,;
    51s1111;.,,1
    65 discussed
    dhove,
    therrn,li
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    5';9

    n'L
    that
    f,i{le.
    ,
    g"lco,tse
    high
    wdt.er
    levr:ls
    drI
    rrilrJiri-:i lrl
    irud,,i{-ii
    i
    e
    ii;:riar-'.j!.::
    ,
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    f
    rorn.
    the iake,
    a
    rtilalivr-.
    ly hiqir
    vOlrr"re ilf
    f
    lOr.in
    Sh*aJ
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    fi+ti:t;j i,,:
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    !r:
    rr:siEt
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    f
    er
    t(-,;npl.rdt
    r;rrl
    int:rr.rJ.,ii
    tft!J
    +1'r,i+i
    i*i::rr,1',
    aqcess
    rotltes
    (by
    connectin,l
    isril,)ted
    nools) ltra',
    atirr'!
    j
    ,in,lii?.:
    1j.;1r,5
    f
    i
    .113r,r3
    !.i-r
    Other
    l()c,lti0n\,
    /rs
    d
    requl'.,
    0ven
    if
    tt,nper,tLiJrr:\
    ,-,r.r:rlr-:*rl
    knt.irrrl
    l,?,.hrl
    j,-.l,ei:
    at
    cer.td,in
    loCal
    ities,
    lhe high
    waLi:r
    woirl..J of
    f
    er e.rpos{i{l
    f
    1:\e.;
    lhi:
    *pf
    iiin.*fi',:'
    mo.
    ing upstr:cani
    Lo
    unaffecLed
    areas or
    Lraveling :Jownrlrear.,
    imLr!
    l'reat
    4t531p:-''
    iion rendered
    waLe!' temperaturcs
    more f
    avorablc. Relolts_
    dl
    _f
    ish
    kl
    l!:1 in
    ',,' .l'
    --djl*T!..r*r.lt-oLthrrjnll.-eff-Lq!llg!1
    o-1
    :l'irdca
    .rat",-,
    u,'ulr-*roii*'
    .
    _,.
    _..__....',*
    -._::.-.
    _-
    :.
    'usly
    raj!_r*even
    f
    or*tJrnpgr'q!ulg:.fl_sj-!.i.vg.f
    orrns,
    rhe
    rri,Jht
    iogicrl
    ly
    ar.,,.Jrir
    '
    that
    Ihe
    abi
    iity
    to detecl
    and
    avoid harrnf
    ul ternper...lures is
    r]nL:
    reisort
    f
    tr
    t.!re
    lack.cf
    s,uch
    rf
    indings. That
    content,ion
    is
    supfior'teri
    rry
    .4ci:.rrllnc
    {
    i976}
    :vrli
    ,
    deterrnined
    tttat stream f
    ishes
    avoided
    potenLially
    leLlral
    Lhcr*al
    e
    f
    f
    lurnls
    ..:..,.
    emanaf ing
    f
    rom
    cool
    inri
    con,Jensers llut
    rapirJly repoouldted
    the
    ,rf
    f
    ect ed str-ei$]
    '
    sect!on,s
    once th€
    flow
    of
    healad
    wdter
    ceasert.
    Strea,n
    f
    islrr:s
    ,rer*',:Js+
    fo:rnrl'i,+
    ,be
    capable of
    travel
    ing great
    distances
    to occupy
    previously
    vac,rt,*rl
    ELr-ean,sqc,
    .tions.tllatwereaffectedbydrought{Larinroretrta]'i959).5'scaptncntit:..
    apparently
    a
    cilrrunon
    sLratngy,tmong
    slrearn
    f
    ishu:,
    provicrlrl
    ,t
    'np,lrl\
    of tvrri,Ji,:r1,...
    deleterious
    ef
    fecIs
    of local
    ly
    extrene o' trnf avoratrle
    envirorr,nent,rl
    car;riir-ir]ni',.,
    rJltereas letltal
    effects
    of
    elevated
    watcr ternpr)raturrs
    or1 fisites
    arr
    rt-.
    laiir,r-,
    1y'
    'eiisy
    Lo'predict, given
    accurate
    tolerance
    dat,r anrl
    consistenI rfspooses
    orr Lhr:
    plrt
    of
    thc f
    ishes,
    srrblrtthal
    ef
    fects
    lenrl
    thern.,e lvcs
    'l
    ess
    r,:.lrJi
    ly
    ti,r
    act-iir'.ttr:
    evaluat
    ion.
    QrranLitative
    ds well
    as
    quaiitativrt
    clranqcs
    dre
    p0siible
    conse-,.:r
    "
    tlugrrees
    of
    lhgrrnal
    enr'l chntont, and these
    cdn occur
    dt ll
    I
    leve
    is
    ,-rf
    or,g.r*i
    jalion
    withirt,the
    f
    ish
    cornmunity,
    In
    Shoal
    Cleek,
    a
    typicel-case
    Lrrrlrrnai
    exposu,'et,
    ',-.'
    -rould
    o(:etlr
    duriilg thc sprinqtirne
    rairty
    te.tton
    whcrr
    ,riter'
    I,;vr,:11
    i!',:
    tli,lit:
    .rrr:l
    ,'.
    cons.Jmer
    pov,rjr
    rlemanrJs
    are
    low. As
    a
    result,
    o
    let,lral
    l4r:rperatrlrrJ
    inr.!-f
    .r.,'*)
    irl
    activities
    ,-
    : ':.
    at that titne
    of
    t€ttf ,
    however,
    arld
    .trt{iiciall.v
    ,:levaLerl
    rrJtcr
    :.
    L.lm-
    -
    perdl.rtres
    on
    the
    spdnnlng
    grountls
    could
    affecL
    r0t)rorJucliv,l
    irr'irc,r!!rrs.
    Ihe
    corr-
    sequences of rJlsrupting or
    rnodiiying
    srtcll
    natural
    f
    urrr.Lion5
    ,tr(r,
    Js
    yet,
    s;)ucg-
    laLive.
    Ihe
    partcity
    of
    gravcl
    srthstrat,es in l.lte
    sLrrty
    arct
    ,;11:.;q*5ts
    tlrat f islrr.:r
    :
    prefe'rrlng
    or
    requiring rocky
    substrates
    for
    spawrrinq
    (
    Ianlc
    5.1)
    ,nay
    rnigrate
    out of
    tfri affected
    a.ea
    prio,
    to
    spawning,
    thus in,rrlverluntl.y,ryoiilrng
    nreas
    t::
    .
    5.
    l0
    l
    l,'
    !:=
    T'
    ltt'
    1...'
    :..:t:
    I
    I
    I
    I
    l,
    :
    t
    T
    l
    |..
    .]
    ,rl
    l'*

    -+l
    -:J
    I
    I
    T
    t
    Sutiject
    to
    thermat
    asJdltir.lns
    in.
    ihe
    sprinr;.'
    :*rcnq
    f
    i.,heq
    !.n*t ril
    s;;.l'd!'i
    ii'i
    r.ne
    ::
    area,
    ,time
    of sparning
    rnay
    be
    advanced anrl
    egtts ant'l
    larv;e +f lh{rse
    f
    i:=i;,:
    \ nh!
    expeiience an accel'eratetl
    ra!e
    of
    developtnent,i'hcn
    er;J+5€rt
    l*
    elsv,tir;'j
    temperatures.
    'Thi.s'can
    be
    perceiverl
    as
    either an enh,lfiieinent
    ur
    o hrnrJrance
    lri
    reproductive
    efforts. An occelerated
    growlh
    r-lte
    Jed
    rj.rr=l'!er
    spa'*ninq
    5eason,
    for
    exampler
    woulrJ of
    fer
    various selective
    advanLages
    to
    earl
    icr
    spawrre,J
    individuals
    compar'ed to conEpecifics
    spawned
    later
    in tir*
    5*ason. 0a
    the
    r:ther
    hand'morerapldt1evelognentduringear}y|iteftaypreYentsynchronizationof
    peak
    plankton
    production
    with the
    onseI
    of cxternal
    feerJing in larvae,
    therrrby
    reducing
    their
    chances
    of survival
    throrrglr Lhe
    criticol
    firsL
    f1.g,1ing
    slages.
    Those
    effects would be
    more
    pronounce(l
    among
    ]arvae Llrat
    are
    displacerl
    downstream
    after hatching and
    utiIize
    aretts other tharr thr:
    innediaLe
    spawninq
    grounds
    for
    feeding
    and
    growth.
    lrr arklilicn, ertqs
    and
    larvae at
    therrnally
    influenced locations
    could he
    erposed to sustained hiqh waler Lemperatures or
    extreme temperature fluctuations
    because,
    rlnlike
    adult
    fishes,
    their
    'l
    irnited
    moti I
    ities
    may
    prevent
    escapernent.
    It
    is
    thus evirJenL
    tlraL llre
    rigi.J rlepenrlency
    of I
    ife
    history
    phenomena
    on the annual therrnal rerline
    'nake
    sl)eci
    f
    ic irnpacls
    of
    unnatural
    temperature
    increases
    diff
    icull
    Lo discern or
    verif-v.
    fuiy
    rrnn,rtur.lI
    temperature
    elevation ma.y aiter the
    capacity of d sl)ecies
    Lo
    survive
    in,rn
    af
    fectecl region by
    mcdif
    ying
    spawning
    tirne
    0rrlocaLi0n,
    nest lui ldinq
    and
    ggarding
    6alivitiei,
    or
    survivcr'ship
    of eqos
    anrl
    I
    arvae
    ,{see
    f
    or exArnp}r',
    Rosenthal and Alderice
    1976).
    .f
    ''-
    rt,"
    I
    5.il

    :i,'
    j:.:.'-
    :.:
    :
    :'
    ,
    LI1[RA[UR[C]T[0
    -
    ::.,'
    Eailey,'R.
    M.,
    J,E.
    Fitch,
    E.S. Haerald,
    t'.
    A.
    Lach6gr,
    C. C.
    -inc-sey,'C.R,,
    Robbins,
    and
    hl.
    G. Scott.
    19i0, ri list
    of
    ccrrynon and
    scicrtifiu ndrncs
    of
    |,sfes
    -f
    rom_the
    Un
    i i.etJ
    St ates and Canada
    ,
    3rd ed,
    trrr. F i
    s,, , Soc
    . Spec
    ,
    Pub I .
    6.
    150
    pp.
    ::
    Balon,_
    E:.K..
    _1975._
    Reproduct!yg
    puj]9r
    9f
    f
    ishes:
    a
    pron,s,rl
    and definjLion.
    .
    J.
    Fish. Res. Board
    Can.
    3?(6):821-864.
    Buslt,
    R.
    H.,
    E. B. l.lelch,
    anrl B.
    I'i.
    Mar.
    19i4. Potential
    effecLs
    of
    thermal
    .
    discha-rges
    on.aquatic
    systems.
    Environ. Sci.
    Tech.
    B{6):561-568.
    '
    .
    Larirnore,
    R. t.l., |,l.
    F. Childers, and
    C.: Heckrotte.
    1959.
    0estruction and
    r€-establ
    ishment
    of
    stream
    fishes
    and
    invertebrates
    affecLed
    by drought.
    Irans,.Am.
    Fish.
    Soc.
    BS(4):26I-285.
    .
    -
    ,
    -
    MacFarlane,
    R.
    ld.
    i976., Fish diversity
    in
    adjacent
    ambient, therrnai, and
    post-thermal
    freshwater
    streams,
    Paqes
    268-271
    in
    G.',''|. Esch and R.'i.
    ,
    MacFarlane,
    eds. Thernal Ecology:ll,'Tech. Infoffi'ation
    Center, Enc'rgy
    Res,
    Oevelop.
    Admin.
    ,:
    '
    l
    Pf'lie$er, til...L.
    1971.
    A
    distributional
    study of
    I'tissouri
    f ishet. tjnjv.
    Kansas
    Publ.,Mus.Nat.Hist.20(3):225-570,:
    rri
    .
    Mi
    ssour
    i
    Dept
    . Conservat
    ion
    .
    343
    pp.
    Rosenthal,.l{.,
    and D. F.''Alderdice.
    1976.
    Sublethal
    effects of
    environrnental
    stressors, natural and
    pol
    lutional, on
    marine fish
    eggs anrl I arvae.
    .1.
    Fish.
    Res.
    Board Can. 33(9):2047-2065.
    5cott,
    H.8.,
    and E. J.
    Crossman.
    19i3. Freshwater
    fistres
    of
    Canada.
    Fjsn.
    Res. Board Can. Bull.
    lB4.
    966
    pp
    Smith, P.
    tJ.
    1979.
    The
    fishes of Il
    l
    inois.
    Univ. ll l.
    Press,
    Urbana.
    3i4
    pp.
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    I.,,.
    l=.'
    !,.:
    i::."
    l,.
    !t,,:
    t-.
    i.i
    l:':'l
    t..t'.
    f"
    t:.
    l=

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